Cambrian arthropods

Hurdia is an extinct genus of hurdiid radiodont that lived 505 million years ago during the Cambrian Period. Fossils have been found in North America, China, and the Czech Republic.

Peytoia is a genus of hurdiid radiodont, an early diverging order of stem-group arthropods, that lived in the Cambrian period, containing two species, Peytoia nathorsti from the Miaolingian of Canada and Peytoia infercambriensis from Poland, dating to Cambrian Stage 3. Its two frontal appendages had long bristle-like spines, it had no fan tail, and its short stalked eyes were behind its large head.

Amplectobelua (meaning "embracing beast") is an extinct genus of late Early Cambrian amplectobeluid radiodont, a group of stem arthropods that mostly lived as free-swimming predators during the first half of the Paleozoic Era.

Kerygmachela kierkegaardi is a kerygmachelid gilled lobopodian from the Cambrian Stage 3 aged Sirius Passet Lagerstätte in northern Greenland. Its anatomy strongly suggests that it, along with its relative Pambdelurion whittingtoni, was a close relative of radiodont (Anomalocaris and relatives) and euarthropods. The generic name "Kerygmachela" derives from the Greek words Kerygma (proclamation) and Chela (claw), in reference to the flamboyant frontal appendages. The specific name, "kierkegaardi" honors Danish philosopher Søren Kierkegaard.

Alalcomenaeus is one of the most widespread and longest-surviving arthropod genera of the Early and Middle Cambrian. Known from over 300 specimens in the Burgess Shale and the Chengjiang biota. It is a member of the family Leanchoiliidae in the group Megacheira.

Canadaspis ("Shield of Canada") is an extinct genus of bivalved Cambrian marine arthropod, known from North America and China. They are thought to have been benthic feeders that moved mainly by walking and possibly used its biramous appendages to stir mud in search of food. They have been placed within the Hymenocarina, which includes other bivalved Cambrian arthropods.

Fuxianhuia is a genus of Lower Cambrian fossil arthropod known from the Chengjiang fauna in China. Its purportedly primitive features have led to it playing a pivotal role in discussions about the euarthropod stem group. Nevertheless, despite being known from many specimens, disputes about its morphology, in particular its head appendages, have made it one of the most controversial of the Chengjiang taxa, and it has been discussed extensively in the context of the arthropod head problem.

300px|left|thumb|Reconstruction of the head region of Cambroraster falcatus. A: Dorsal view, B: Ventral view, Ey: Eye, Fa: Frontal appendage, He:H-element, Bp: Bilobate posterior region, Lp: Posterolateral process, Oc: Oral cone, Pe: P-element, Pn: P-element neck Cambroraster is an extinct monotypic genus of hurdiid radiodont, dating to the middle Cambrian, and represented by the single formally described species Cambroraster falcatus. Hundreds of specimens were found in the Burgess Shale, and described in 2019. A large animal (for its era) at up to (but not as long as Titanokorys at ), it is

Yohoia is an extinct genus of fossil megacheiran arthropod from the Cambrian period. The type species, Yohoia tenuis, has been found in the Burgess Shale formation of British Columbia. 711 specimens of Yohoia are known from the Greater Phyllopod bed, where they comprise 1.35% of the community. In 2015, Conway Morris et al. reported another species, Y. utahana, from the Marjum Formation, Utah.

Utaurora is an extinct genus of opabiniid, which were stem-arthropods closely related to true arthropods and radiodonts; the type species is U. comosa. The animal's fossils come from the Cambrian of Utah. This genus is the only other known unquestionable opabiniid, with the other being Opabinia itself. There are other animals like Myoscolex and Mieridduryn that could be opabiniids, but the classification of those two genera is still debated.

Waptia is an extinct genus of marine arthropod from the Middle Cambrian of North America. It grew to a length of , and had a large bivalved carapace and a segmented body terminating into a pair of tail flaps. It was an active swimmer and likely a predator of soft-bodied prey. It is also one of the oldest animals with direct evidence of brood care. Waptia fieldensis is the only species classified under the genus Waptia, and is known from the Burgess Shale Lagerstätte of British Columbia, Canada. Specimens of Waptia are also known from the Spence Shale of Utah, United States.

Titanokorys is a genus of extinct hurdiid (peytoiid) radiodont (a grouping of primitive stem arthropods which lived during the early Paleozoic) that lived during the Miaolingian epoch of the middle Cambrian. It is the largest member of its family from the Cambrian, with an estimated body length of around long, making it one of the largest animals of the time. It bears a resemblance to the related, and contemporary, genus Cambroraster. Fossils of T. gainesi were first found within the Marble Canyon locality of the Burgess Shale in 2018, however the fossils were not named until 2021 because they

Parapeytoia is a genus of Cambrian arthropod. The type and only described species is Parapeytoia yunnanensis, which lived over 518 million years ago (Cambrian Stage 3) in the Maotianshan shales of Yunnan, China. Unidentified fossils from the same genus have also been discovered from the nearby Wulongqing Formation (Cambrian Stage 4).

Mollisonia is an extinct genus of Cambrian marine arthropod. Four species have been described from North America and China. Studies suggest it is a basal member of Chelicerata, a group which includes horseshoe crabs and arachnids.

Tamisiocarididae is a family of radiodonts, extinct marine animals related to arthropods, that bore finely spined appendages that were presumably used in filter-feeding. When first discovered, the clade was named Cetiocaridae after a speculative evolution artwork, Bearded Ceticaris by John Meszaros, that depicted a hypothetical filter-feeding radiodont at a time before any were known to exist. However, the family name was not valid according to the International Code of Zoological Nomenclature, as no real genus named "Cetiocaris" exists, and in 2019 it was formally replaced by the name Tamisio

thumb|270x270px|Agnostus sp. from the Middle Cambrian of Couloma, Hérault, France. Max Rouger Collection.

Kylinxia is a genus of extinct arthropod described in 2020. It was described from six specimens discovered in Yu'anshan Formation (Maotianshan Shales) in southern China. The specimens are assigned to one species Kylinxia zhangi. Dated to 518 million years, the fossils falls under the Cambrian period. Announcing the discovery on 4 November 2020 at a press conference, Zeng Han of the Nanjing Institute of Geology and Paleontology, said that the animal "bridges the evolutionary gap from Anomalocaris to true arthropods and forms a key ‘missing link’ in the origin of arthropods," which was "predicte

Sarotrocercus is a small Cambrian arthropod known from Burgess Shale, reaching a centimetre or two in length (). Sarotrocercus is only known from 7 specimens. It may lie in the arthropod crown group, and a recent study has revised some points of its original description.

Burgessia is a genus of arthropod known from the mid-Cambrian aged Burgess Shale of British Columbia, Canada. It is relatively abundant, with over 1,700 specimens having been collected.

Habelia is a genus of extinct arthropod from the Middle Cambrian, thought to be one of the earliest known relatives of chelicerates. Its fossils have been found in the Burgess Shale in British Columbia, Canada. Fifty-four specimens of Habelia are known from the Greater Phyllopod bed, where they comprise 0.1% of the community.

Isoxys (meaning "equal surfaces") is a genus of extinct bivalved Cambrian arthropod; the various species of which are thought to have been freely swimming predators. It had a pair of large spherical eyes (which are the most commonly preserved feature of the soft-bodied anatomy), and two large frontal appendages used to grasp prey.

Pambdelurion is an extinct genus of panarthropod from the Cambrian aged Sirius Passet site in northern Greenland. Like the morphologically similar Kerygmachela from the same locality, Pambdelurion is thought to be closely related to arthropods, combining characteristics of "lobopodians" with those of primitive arthropods.

Helmetia is an extinct genus of arthropods from the middle Cambrian (Wuliuan). Fossils of the type species Helmetia expansa have been found in the Burgess Shale of Canada. It is characterized by a pair of spines in front of its head. It is classified under Conciliterga, a group of trilobitomorph artiopod.

Misszhouia is a genus of small to average sized (up to long) marine trilobite-like arthropods within the Naraoiidae family, that lived during the early Cambrian period. The species are M. longicaudata, from the Maotianshan Shales, described in 1985, and M. canadensis, from the Burgess Shale and described in 2018, although later species may belongs to genus Naraoia instead.

Laminacaris is a genus of extinct stem-group arthropods (Radiodonta) that lived during the Cambrian period. It is monotypic with a single species Laminacaris chimera, the fossil of which was described from the Chengjiang biota of China in 2018. Around the same time, two specimens that were similar or of the same species were discovered at the Kinzers Formation in Pennsylvania, USA. The first specimens from China were three frontal appendages, without the other body parts.

Haikoucaris is a genus of megacheiran arthropod that contains the single species Haikoucaris ercaiensis. It was discovered in the Cambrian Chengjiang biota of China.

Hymenocaris is a genus of Cambrian marine arthropod. The type species, Hymenocaris vermicaudata is known from the Upper Cambrian (Furongian) Ffestiniog Formation, in North Wales, United Kingdom.

Skania is a Cambrian fossil arthropod. The type species, S. fragillis, is known from the Burgess Shale of British Columbia, Canada. A second possible species "S." sundbergi is known from the Kaili Formation of China, but its placement within the genus has been questioned.

species of Cambrian arthropods

Kootenichela deppi is an extinct arthropod described from the Middle Cambrian of the Kootenay National Park, Canada. It is originally considered to be a member of "great appendage arthropods", although subsequent studies questioned its affinity. Kootenichela appears to be the sister taxon of Worthenella, from cladistic analysis.

Hymenocarina is an order of extinct marine arthropods known from the Cambrian. They possess bivalved carapaces, typically with exposed posteriors. Members of the group are morphologically diverse and had a variety of ecologies, including as filter feeders and as predators. Recent research has generally considered them to be stem or crown group members of Mandibulata, due to the presence of mandibles in well-preserved species.

Paranomalocaris is a genus of radiodont recovered from the Cambrian Stage 4 aged Wulongqing Formation, eastern Yunnan. It contains two species, Paranomalocaris multisegmentalis and P. simplex. It is only known from its frontal appendage.

Erratus is an extinct genus of marine arthropod from the Cambrian of China. Its type and only species is Erratus sperare. Erratus is likely one of the most basal known arthropods, and its discovery has helped scientists understand the early evolution of arthropod trunk appendages. Some of the stem-arthropods like radiodonts did not have legs, instead they had flap like appendages that helped them swim. Erratus on the other hand had not only flaps but also a set of primitive legs. It also supported the theory that the gills of aquatic arthropods probably evolved into the wings and lungs of terr

left|thumb|Life restoration

Chengjiangocaris is an extinct genus of fuxianhuiid arthropod known from the Cambrian of South China. It contains two species, C. longiformis which was described in 1991. C. kunmingensis was described in 2013 by Javier Ortega-Hernández and colleagues. One specimen of C. kunmingensis shows detailed evidence of a nervous system. The nervous system of the chengjiangocaris is and has always been very complex to understand but a recent discovery of the ladder like ventral nerval cords and segmental ganglia of the related chengjiangocaris together with the brain provides the most comprehensive recon

Protichnites is an ichnogenus of trace fossil consisting of the imprints made by the walking activity of certain arthropods. It consists of two rows of tracks and a medial furrow between the two rows. This furrow, which may be broken, set at an angle, and of varying width and depth, is thought to be the result of the tail region contacting the substrate.

Balhuticaris is a genus of extinct bivalved (referring to the carapace) hymenocarine arthropod that lived in the Cambrian aged Burgess Shale in what is now British Columbia around 506 million years ago. This extremely multisegmented (with over 100 segments) arthropod is the largest member of the group, and it was even one of the largest animals of the Cambrian, with individuals reaching lengths of 245 mm (9 in). Fossils of this animal suggests that gigantism occurred in more groups of Arthropoda than had been previously thought. It also presents the possibility that bivalved arthropo

Occacaris is an extinct nektonic predatory arthropod from the Lower Cambrian Maotianshan shale Lagerstätte, known from three species. It bears a superficial resemblance to the Cambrian arthropod, Canadaspis, though, was much smaller, and had a pair of "great appendages", with which it may have grasped prey. It was originally considered to belong to Megacheira, however it is questioned in later study.

Jugatacaris is an extinct genus of bivalved arthropod known from a single species, Jugatacaris agilis found in the Cambrian Stage 3 aged Chengjiang biota of Yunnan, China. The carapace is around in length, with a pronounced ridge at the top of the carapace separating the two valves, which formed a fin-like structure raised above the carapace. The head has a pair of stalked eyes, as well as a dumbbell shaped medial eye between them. The head also bore a pair of mandibles as well as at least one and possibly two pairs of antennae. The trunk had up to 65 segments, each with biramous (two-branched

Pahvantia is an extinct genus of hurdiid radiodont from the Cambrian period. It is known by a single species, Pahvantia hastata, described from Wheeler Shale and Marjum Formation in Utah. Although it was once considered as a filter feeder based on the large number of putative setae on the frontal appendage, these structures were later interpreted as fossil material of the "trunk" section.

Forfexicaris valida is a species of Lower Cambrian arthropod, the only species in the family Forfexicarididae. It is known from only two specimens from the Maotianshan shale ''''.

Molaria is a genus of Cambrian arthropod, the type species M. spinifera is known from the Middle Cambrian Burgess Shale. 144 specimens of Molaria are known from the Greater Phyllopod bed, where they comprise 0.27% of the community. A second species M. steini was described from the Sirius Passet in Greenland in 2017.

Yawunik is an extinct genus of Cambrian megacheiran ("Great appendage" arthropod) known from the Burgess Shale in Canada ( Marble Canyon locality). The type species has been named Yawunik kootenayi after the Kootenay, both a geographic area (and National Park, where the fossil was found) and North American First Nation, also known as the Ktunaxa. The genus name is derived from , the name of a primordial sea monster in Ktunaxa mythology. The fossil dates back to 508 million years ago.

Echidnacaris briggsi is an extinct species of radiodont known from the Cambrian Stage 4 aged Emu Bay Shale of Australia. Formerly referred to as "Anomalocaris" briggsi, it was placed in the new monotypic genus Echidnacaris in 2023. It is only distantly related to true Anomalocaris, and is instead placed in the family Tamisiocarididae.