Category
page 1Carboniferous first appearances

Orthoptera
Orthoptera () is an order of insects that comprises the grasshoppers, locusts, and crickets, including closely related insects, such as the bush crickets or katydids and wētā. The order is subdivided into two suborders: Caelifera – grasshoppers, locusts, and close relatives; and Ensifera – crickets and close relatives.

Pulmonata
Pulmonata or pulmonates is an informal group (previously an order, and before that, a subclass) of snails and slugs characterized by the ability to breathe air, by virtue of having a pallial lung instead of a gill, or gills. The group includes many land and freshwater families, and several marine families.

Palaeoptera
The name Palaeoptera (from Greek ( 'old') + ( 'wing')) has been traditionally applied to those ancestral groups of winged insects (most of them extinct) that lacked the ability to fold the wings back over the abdomen as characterizes the Neoptera. The Diaphanopterodea, which are palaeopteran insects, had independently and uniquely evolved a different wing-folding mechanism. Both mayflies and dragonflies lack any of the smell centers in their brain found in Neoptera. Their midgut epithelium has a dual origin: the anterior and posterior regions develop through cellular differentiation, while the

Coleoidea
Coleoidea or Dibranchiata is one of the two subclasses of cephalopod molluscs containing all the various taxa popularly thought of as "soft-bodied" or "shell-less" (i.e. octopus, squid and cuttlefish). Unlike its sister groups, the shelled Ammonoidea and Nautiloidea, the coleoids have at most an internal shell called cuttlebone or gladius that is used for buoyancy or as muscle attachment. Some species, notably the incirrate octopuses (including commonly known varieties living in the shallows), have lost their internal shell altogether, while in some it has been replaced by a chitinous support

Dictyoptera
thumb|upright|Termite queen with soldiers
Dictyoptera (from Greek δίκτυον diktyon "net" and πτερόν pteron "wing") is an insect superorder that includes two extant orders of polyneopterous insects: the order Blattodea (termites and cockroaches together) and the order Mantodea (mantises). All modern Dictyoptera have short ovipositors and typically lay oothecae. The oldest fossils of Dictyoptera from the Late Carboniferous, referred to as "roachoids" have long ovipositors and did not lay oothecae. The oldest modern oothecae-laying dictyopterans date to the Late Triassic.
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==Classification

Opisthobranchia
Opisthobranchs () is a now informal name for a large and diverse group of specialized complex gastropods which used to be united in the subclass Opisthobranchia. That taxon is no longer considered to represent a monophyletic grouping.
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Calamites
Calamites is a genus of extinct arborescent (tree-like) horsetails to which the modern horsetails (genus Equisetum) are closely related. Unlike their herbaceous modern cousins, these plants were medium-sized trees, growing to heights of . They were components of the understories of coal swamps of the Carboniferous Period (around ).

Caenogastropoda
Caenogastropoda is a taxonomic subclass of molluscs in the class Gastropoda. It is a large diverse group which are mostly sea snails and other marine gastropod mollusks, but also includes some freshwater snails and some land snails. The subclass is the most diverse and ecologically successful of the gastropods.
Basommatophora
Basommatophora was a term that was previously used as a taxonomic informal group, a group of snails within the informal group Pulmonata, the air-breathing slugs and snails. According to the taxonomy of the Gastropoda (Bouchet & Rocroi, 2005), whenever monophyly has not been tested, or where a traditional taxon of gastropods has now been discovered to be paraphyletic or polyphyletic, the term "group" or "informal group" was used.

Lepidodendrales
Lepidodendrales (from the Greek for "scale tree") or arborescent lycophytes are an extinct order of primitive, vascular, heterosporous, arborescent (tree-like) plants belonging to Lycopodiopsida. Members of Lepidodendrales are the best understood of the fossil lycopsids due to the vast diversity of Lepidodendrales specimens and the diversity in which they were preserved; the extensive distribution of Lepidodendrales specimens as well as their well-preservedness lends paleobotanists exceptionally detailed knowledge of the coal-swamp giants’ reproductive biology, vegetative development, and role

Sigillaria
Sigillaria is a genus of extinct, spore-bearing, arborescent lycophyte, known from the Carboniferous and Permian periods. It is related to the more famous Lepidodendron, and more distantly to modern quillworts.

Calamitaceae
Calamitaceae is an extinct family of equisetalean plants related to the modern horsetails, known from the Carboniferous and Permian periods. Some members of this family like Arthropitys attained tree-like stature, with heights over , with extensive underground rhizomes. They were largely found in wetland environments.

Hybodontiformes
Hybodontiformes, commonly called hybodonts, are an extinct group of shark-like cartilaginous fish (chondrichthyans) which existed from the late Devonian to the Late Cretaceous. Hybodonts share a close common ancestry with modern sharks and rays (Neoselachii) as part of the clade Euselachii. They are distinguished from other chondrichthyans by their distinctive fin spines and cephalic spines present on the heads of males. An ecologically diverse group, they were abundant in marine and freshwater environments during the late Paleozoic and early Mesozoic, but were rare in open marine environments

Colosteidae
Colosteidae is a family of stegocephalians (stem-group tetrapods) that lived in the Carboniferous period. They possessed a variety of characteristics from different tetrapod or stem-tetrapod groups, which made them historically difficult to classify. They are now considered to be part of a lineage intermediate between the earliest Devonian terrestrial vertebrates (such as Ichthyostega), and the different groups ancestral to all modern tetrapods, such as temnospondyls (probably ancestral to modern amphibians) and reptiliomorphs (ancestral to amniotes such as mammals, reptiles, and birds).
Miomoptera
Miomoptera is an extinct order of insects. Although it is thought to be a common ancestor of all holometabolous insects, because no smooth transition between Miomoptera and other holometabolous insect orders is known, it is considered to be in a separate order unto itself.
Argulidae
The family Argulidae, whose members are commonly known as carp lice or fish lice, (not to be confused with Caligidae, also commonly referred to as sea lice) are parasitic crustaceans in the class Ichthyostraca. It is the only family in the monotypic subclass Branchiura and the order Arguloida, although a second family, Dipteropeltidae, has been proposed.
Annularia
Annularia is a form taxon, applied to fossil foliage belonging to extinct plants of the genus Calamites in the order Equisetales.

Chiton
genus of molluscs
Brachyphyllum
Brachyphyllum (meaning "short leaf") is a form genus of fossil coniferous plant foliage. Plants of the genus have been variously assigned to several different conifer groups including Araucariaceae and Cheirolepidiaceae. They are known from around the globe from the Late Carboniferous to the Late Cretaceous periods.
Miliolida
The Miliolida are an order of foraminifera with calcareous, porcelacous tests that are imperforate and commonly have a pseudochitinous lining. Tests are composed of randomly oriented calcite needles that have a high proportion of magnesium along with organic material. Tests lack pores and generally have multiple chambers.
Pygocephalomorpha
The order Pygocephalomorpha is an extinct group of peracarid crustaceans. Pygocephalomorpha appeared in the Late Devonian, were abundant from the Carboniferous era until their extinction in the Early Permian era.
Blattoptera
"Roachoids", also known as "Roachids", "Blattoids" or Eoblattodea, are members of the stem group of Dictyoptera (the group containing modern cockroaches, termites and praying mantises). They generally resemble cockroaches, but most members, unlike modern dictyopterans, have generally long external ovipositors, and are thought not to have laid ootheca like modern dictyopterans.
Callistophytales
Callistophytales is an extinct order of spermatophytes (seed plants) which lived from the Pennsylvanian (Late Carboniferous) to Permian periods. They were mainly scrambling and lianescent (vine-like) plants found in the wetland "coal swamps" of Euramerica and Cathaysia. Like many other early spermatophytes, they could be described as "seed ferns", combining ovule-based reproduction with pinnate leaves superficially similar to modern ferns.
Medullosales
The Medullosales is an extinct order of pteridospermous seed plants characterised by large ovules with circular cross-section and a vascularised nucellus, complex pollen-organs, stems and rachides with a dissected stele, and frond-like leaves. Their nearest still-living relatives are the cycads.
Alethopteris
Alethopteris is a prehistoric plant genus of fossil pteridospermatophytes (seed ferns) that developed in the Pennsylvanian (around ).
thumb|left|Alethopteris florentina De Stefani, 1901, Natural History Museum University of Pisa
thumb|Alethopteris, at the State Museum of Pennsylvania in Harrisburg.|left
thumb|Alethopteris sp.
Phlegethontiidae
Phlegethontiidae is a family of extinct aistopod tetrapodomorphs including the genera Phlegethontia and Sillerpeton.