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Cenomanian dinosaurs

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Spinosaurus
Spinosaurus (; ) is a genus of large spinosaurid theropod dinosaurs that lived in what is now North Africa during the Cenomanian stage of the Late Cretaceous period, about 100 to 94 million years ago. The genus was known first from Egyptian remains discovered in 1912 and described by German palaeontologist Ernst Stromer in 1915. The original remains were destroyed in World War II, but additional material came to light in the early 21st century. It is unclear whether one or two species are represented in the fossils reported in the scientific literature. The type species, S. aegyptiacus, is mai
Carcharodontosaurus
Carcharodontosaurus (; "shark-toothed lizard") is a genus of large theropod dinosaur that lived in Northwest Africa from about 100 to 94 million years ago during the Cenomanian stage of the Cretaceous. The taxon was first described in 1925 by French paleontologists Charles Depéret and Justin Savornin as Megalosaurus saharicus, based on two fossil teeth discovered in Algeria, which are now lost. A partial skeleton was discovered in Egypt as early as 1914 by crews of German paleontologist Ernst Stromer, although he did not report the find until 1931. Based on this specimen, together with the tee
Giganotosaurus
Giganotosaurus ( ) is a genus of large theropod dinosaur that lived in what is now Argentina, during the early Cenomanian age of the Late Cretaceous period, approximately 99.6 to 95 million years ago. The holotype specimen was discovered in the Candeleros Formation of Patagonia in 1993 and is almost 70% complete. The animal was named Giganotosaurus carolinii in 1995; the genus name translates to "giant southern lizard", and the specific name honors the discoverer, Ruben Carolini. A dentary bone, a tooth, and some tracks, discovered before the holotype, were later assigned to this animal. The g
Aegyptosaurus
Aegyptosaurus (meaning 'Egypt's lizard') is a genus of sauropod dinosaur discovered in Egypt, that lived in what is now Africa, around 95 million years ago, during the Late Cretaceous Period (Cenomanian faunal stage).
Buitreraptor
Buitreraptor (meaning "La Buitrera seizer") is a genus of unenlagiine theropod dinosaurs that lived during the Late Cretaceous of Argentina at the Candeleros Formation. Buitreraptor was described in 2005 and the type species is Buitreraptor gonzalezorum. It was rooster-sized and had a very elongated head with many small teeth.
Muttaburrasaurus
Muttaburrasaurus was a genus of herbivorous iguanodontian ornithopod dinosaur that lived in what is now north-eastern Australia sometime between 112 and 103 million years ago during the early Cretaceous period. It has been recovered in some analyses as a member of the iguanodontian clade Rhabdodontomorpha, or alternately as a member of Elasmaria. After Kunbarrasaurus, it is Australia's most completely known dinosaur from skeletal remains. It was named after Muttaburra, the site in Queensland, Australia, where it was found. The dinosaur was selected from twelve candidates to become the official
Nodosaurus
Nodosaurus (meaning 'knobbed lizard') is a genus of herbivorous nodosaurid ankylosaurian dinosaur from the Late Cretaceous, the fossils of which are found exclusively in the Frontier Formation in Wyoming.
Paralititan
Paralititan (meaning "tidal giant") is a genus of sauropod dinosaur that lived in present-day Egypt during the Late Cretaceous period. It was described by American paleontologist Joshua B. Smith and colleagues in 2001. The genus contains a single species, Paralititan stromeri, named based on a fragmentary skeleton including vertebrae and limb bones. These fossils were unearthed by an American expedition to the Bahariya Oasis in western Egypt in rock layers of the Bahariya Formation. This formation dates to the Cenomanian stage of the Late Cretaceous, which lasted . An incomplete (back) vertebr
Rugops
Rugops (meaning 'wrinkle face') is a monospecific genus of basal abelisaurid theropod dinosaur that lived during the Late Cretaceous period (Cenomanian stage, approximately 95 million years ago) in what is now the Echkar Formation in Niger. The type and only species, Rugops primus, is known only from a partial skull. It was named and described in 2004 by Paul Sereno, Jeffery Wilson and Jack Conrad. Rugops has an estimated length of and weight of . The top of its skull bears several pits which correlates with overlaying scale and the front of the snout would have had an armour-like dermis.
Australovenator wintonensis
Australovenator (meaning "southern hunter") is a genus of megaraptoran theropod dinosaur from the Late Cretaceous Winton Formation (Cenomanian, 95 million years ago) of Australia. Some specimens from the Albian-aged Eumeralla Formation and the Wonthaggi Formation may belong to Australovenator. It is known from partial cranial and postcranial remains, which were described in 2009 by Scott Hocknull and colleagues, although additional descriptions and analyses continue to be published. It is the most complete predatory dinosaur discovered in Australia. It has been suggested that Australovenator i
Oxalaia
Oxalaia (in reference to the African deity Oxalá) is a genus of spinosaurid dinosaur that lived in what is now the Northeast Region of Brazil during the Cenomanian stage of the Late Cretaceous period, sometime between 100.5 and 93.9 million years ago. Its fossils were found in 1999 on Cajual Island in the rocks of the Alcântara Formation, which is known for its abundance of fragmentary, isolated fossil specimens as well as the Itapecuru Formation. The remains of Oxalaia were described in 2011 by Brazilian palaeontologist Alexander Kellner and colleagues, who assigned the specimens to a new gen
Wintonotitan
Wintonotitan (meaning "Winton titan") is a genus of titanosauriform dinosaur from the Cenomanian-aged (Late Cretaceous) Winton Formation of Australia. It is known from partial postcranial remains.
Andesaurus
Andesaurus ( ; "Andes lizard") is a genus of basal titanosaurian sauropod dinosaur which existed during the middle of the Cretaceous Period in South America. Like most sauropods, it would have had a small head on the end of a long neck and an equally long tail.
Archaeornithomimus
Archaeornithomimus (meaning "ancient bird mimic") is a genus of ornithomimosaurian theropod dinosaur that lived during the Late Cretaceous period (around 96 million years ago) in the Iren Dabasu Formation of Inner Mongolia, China.
Bahariasaurus
Bahariasaurus (meaning "Bahariya lizard") is an enigmatic genus of large theropod dinosaur. The genus contains a single species, Bahariasaurus ingens, which was found in North African rock layers dating to the Cenomanian age of the Late Cretaceous. The only fossils confidently assigned to Bahariasaurus were found in the Bahariya Formation of the Bahariya Oasis in Egypt by Ernst Stromer. This material was destroyed during a World War II bombing raid, with the same raid also destroying the holotypes of Spinosaurus, Aegyptosaurus, and other animals found in the Bahariya Formation.
Oryctodromeus
Oryctodromeus (from Ancient Greek ὀρυκτήρ (oruktḗs), meaning "digger", and δρομαῖος (dromaîos), meaning "runner") is an extinct genus of small orodromine thescelosaurid dinosaur. Fossils are known from the Late Cretaceous Blackleaf Formation of southwestern Montana and the Wayan Formation of southeastern Idaho, USA, both of the Cenomanian stage, roughly 105-96 million years ago. A member of the small, presumably fast-running herbivorous family Thescelosauridae, Oryctodromeus is the first non-avian dinosaur published that shows evidence of burrowing behavior.
Eolambia
Eolambia (meaning "dawn lambeosaurine") is a genus of herbivorous hadrosauroid dinosaur from the early Late Cretaceous of the United States. It contains a single species, E. caroljonesa, named by paleontologist James Kirkland in 1998. The type specimen of Eolambia was discovered by Carole and Ramal Jones in 1993; the species name honors Carole. Since then, hundreds of bones have been discovered from both adults and juveniles, representing nearly every element of the skeleton. All of the specimens have thus far been found in Emery County, Utah, in a layer of rock known as the Mussentuchit Membe
Rebbachisaurus
Rebbachisaurus (meaning "Aït Rebbach lizard") is a genus of sauropod dinosaur that lived in what is now Africa during the Late Cretaceous period, between 99 and 97 million years ago. The only valid species is R. garasbae. However, two other species have been assigned to the genus; R. tessonei, now Limaysaurus tessonei, and the nomen dubium R. tamesnensis. Known from a single, incomplete skeleton, much of R. garasbae's anatomy is unknown. The skeleton, discovered in 1948, was unearthed in the Errachidia Province of Morocco from strata of the Kem Kem Beds.
Deltadromeus
Deltadromeus (meaning "river delta or change runner") is an extinct genus of controversial theropod dinosaur that lived in present-day Morocco during the mid-Cretaceous period. It was described by American paleontologist Paul Sereno and colleagues in 1996. The genus contains a single species, D. agilis, named based on an incomplete postcranial skeleton, the holotype specimen. However, some fossils from the Bahariya Formation of Egypt that were formerly referred to the theropod Bahariasaurus have been suggested to belong to Deltadromeus. The holotype specimen of Deltadromeus was unearthed by a
Skorpiovenator
Skorpiovenator ("scorpion hunter") is a genus of abelisaurid theropod dinosaur from the Late Cretaceous (Cenomanian to Turonian) Huincul Formation of Argentina. The sole species of Skorpiovenator, S. bustingorryi, was named in honour of Manuel Bustingorry, the late owner of the farm on which the type specimen was discovered. Formally described in 2009, the type specimen is one of the most complete and informative abelisaurids yet known, described from a nearly complete and articulated skeleton. A tibia fragment was assigned to Skorpiovenator in 2022.
Animantarx
Animantarx ( ; meaning 'living citadel') is a genus of nodosaurid ankylosaurian dinosaur from the Early and Late Cretaceous of western North America. Like other nodosaurs, it would have been a slow-moving quadrupedal herbivore covered in heavy armor scutes, but without a tail club. The skull measures approximately 25 cm (10 inches) in length, suggesting the animal as a whole was no more than 3 meters (10 feet) long.
Aniksosaurus darwini
Aniksosaurus (meaning "spring lizard", from Modern Greek Άνοιξη, "Spring", referring to the fact it was found on 21 September 1995, the onset of Spring on the Southern Hemisphere) is a genus of avetheropod dinosaur from what is now Chubut Province, Argentina. It lived during the Cenomanian to Turonian of the Cretaceous period, between 96-91 million years ago. The type species, Aniksosaurus darwini, was formally described from the Bajo Barreal Formation of the Golfo San Jorge Basin by Rubén Dario Martínez and Fernando Emilio Novas in 2006; the name was first coined in 1995 and reported in the l
Xenotarsosaurus
Xenotarsosaurus is a genus of abelisaurid theropod dinosaur that lived during the Late Cretaceous of Argentina.
Siats
Siats (/see-ats/) is an extinct genus of large theropod dinosaurs known from the Late Cretaceous Cedar Mountain Formation (Mussentuchit Member) of Utah, United States. The genus contains a single species, Siats meekerorum. It was initially classified as a megaraptoran, a clade of large theropods with controversial relationships. Alternative positions within the Neovenatoridae, Allosauroidea, and Tyrannosauroidea have also been proposed.
Sigilmassasaurus
Sigilmassasaurus ( ; "Sijilmassa lizard") is a controversial genus of spinosaurid dinosaur that lived approximately 100 to 94 million years ago during the Late Cretaceous Period in what is now northern Africa. Named in 1996 by Canadian paleontologist Dale Russell, it contains a single species, Sigilmassasaurus brevicollis. The identity of the genus has been debated by scientists, with some considering its fossils to represent material from the closely related species Spinosaurus aegyptiacus, while others have classified it as a separate taxon, forming the clade Spinosaurini with Spinosaurus as
Asiaceratops
Asiaceratops (meaning "Asian horned face") is a genus of herbivorous ceratopsian dinosaur. It lived during the Early-Late Cretaceous. The type species, A. salsopaludalis is known from Uzbekistan, while A. sulcidens is known from China and Mongolia.
Gilmoreosaurus
Gilmoreosaurus (meaning "Charles Whitney Gilmore's lizard") is a genus of hadrosauromorphan dinosaur from the Late Cretaceous of Asia. The type species is Gilmoreosaurus mongoliensis, known from the Iren Dabasu Formation of Inner Mongolia, dating to 96 million years ago. Additional specimens have been described as distinct species, including G. atavus from the Khodzhakul Formation of Uzbekistan, and G. arkhangelskyi from the Bissekty Formation, also of Uzbekistan. However, these are based on very fragmentary remains, and their classification is dubious. An additional species, G. kysylkumense,
Ekrixinatosaurus
Ekrixinatosaurus ('explosion-born reptile') is a genus of abelisaurid theropod which lived approximately 99 to 97 million years ago during the Late Cretaceous period. Its fossils have been found in Argentina. Only one species is currently recognized, Ekrixinatosaurus novasi, from which the specific name honors Dr. Fernando Novas for his contributions to the study of abelisaurid theropods, while the genus name refers to the dynamiting of the holotype specimen. It was a large abelisaurid, measuring between in length and weighing .
Ilokelesia
Ilokelesia is an extinct genus of abelisaurid theropod dinosaurs, preserved in the layers of the earliest Late Cretaceous of the Huincul Formation (Neuquén Group) in Neuquén Province, Argentina. The only known specimen, consisting of fragmentary elements of the skull and skeleton, was described by Rodolfo Coria and Leonardo Salgado in late 1998.
Jakapil
Jakapil (meaning "shield bearer" in Puelchean) is a genus of basal thyreophoran dinosaur from the Late Cretaceous (Cenomanian-aged) Candeleros Formation of Argentina. The type species is Jakapil kaniukura.
Limaysaurus
Limaysaurus ("Limay lizard") is a genus of rebbachisaurid sauropod dinosaurs which lived during the late Cretaceous period, about 99.7 to 94 million years ago, in the Cenomanian, in what is now South America (northwestern Patagonia). It is represented by a single species, L. tessonei.
Cedarpelta
Cedarpelta is an extinct genus of basal ankylosaurid dinosaur from Utah that lived during the Late Cretaceous period (Cenomanian to lower Turonian stage, 98.2 to 93 Ma) in what is now the Mussentuchit Member of the Cedar Mountain Formation. The type and only species, Cedarpelta bilbeyhallorum, is known from multiple specimens including partial skulls and postcranial material. It was named in 2001 by Kenneth Carpenter, James Kirkland, Don Burge, and John Bird. Cedarpelta has an estimated length of 7 metres (23 feet) and weight of 5 tonnes (11,023 lbs). The skull of Cedarpelta lacks extensi
Sauroniops
Sauroniops is a controversial genus of carnivorous carcharodontosaurid theropod dinosaurs known from the Late Cretaceous (Cenomanian stage) of Morocco. It is known from the Gara Sbaa Formation of the Kem Kem beds and contains a single species, S. pachytholus.
Crichtonsaurus
Crichtonsaurus (meaning "Crichton's lizard") is a dubious genus of herbivorous ankylosaurid dinosaur that lived during the Late Cretaceous in what is now China. It was named after Michael Crichton, the author of the dinosaur novel Jurassic Park. A sister taxon was discovered, C. benxiensis, which is now identified as a separate genus.
Rapator
Rapator is a genus of theropod dinosaur from the Griman Creek Formation of New South Wales, Australia, dating to the Cenomanian age of the Cretaceous period. It contains only the type species, Rapator ornitholestoides, which was originally named by Friedrich von Huene in 1932.
Bicentenaria
Bicentenaria (meaning "two hundred year anniversary"; named after the 200th anniversary of the 1810 May Revolution in Argentina) is an extinct genus of carnivorous coelurosaurian theropods which lived during the early Late Cretaceous (Cenomanian) Candeleros Formation of Argentina. It contains the type species, B. argentina.
Walgettosuchus
Walgettosuchus (meaning "Walgett crocodile") is a dubious or possibly invalid genus of extinct tetanuran theropod dinosaur that lived in Australia during the Late Cretaceous (Cenomanian). It is known from a single caudal vertebra.
Rayososaurus
Rayososaurus is an extinct genus of sauropod dinosaur in the family Rebbachisauridae. Rayososaurus was named by Argentinian paleontologist José Bonaparte in 1996. Its type and only accepted species is Rayososaurus agrioensis. The species Limaysaurus tessonei was at one point included in Rayososaurus as Rayososaurus tessonei.
Zhejiangosaurus
Zhejiangosaurus (meaning "Zhejiang lizard") is an extinct genus of ankylosaurian dinosaur from the Upper Cretaceous (Cenomanian stage) of Zhejiang, eastern China. It was first named by a group of Chinese authors Lü Junchang, Jin Xingsheng, Sheng Yiming and Li Yihong in 2007 and the type species is Zhejiangosaurus lishuiensis ("from Lishui", where the fossil was found). It has no diagnostic features, and thus is a nomen dubium.
Erliansaurus
Erliansaurus (meaning "Erlian lizard") is a genus of therizinosaurian theropod dinosaur that lived in Asia during the Cenomanian stage of the Late Cretaceous period in what is now Nei Mongol, Iren Dabasu Formation.
Ulughbegsaurus
Ulughbegsaurus (meaning "Ulugh Beg's lizard") is an extinct genus of carcharodontosaurid theropod dinosaurs from the Late Cretaceous Bissekty Formation (Turonian age) and Khodzhakul Formation (Cenomanian age) of Uzbekistan. The genus contains a single species, Ulughbegsaurus uzbekistanensis, known from multiple maxilla fragments.
Silvisaurus
Silvisaurus, from the Latin silva "woodland" and Greek sauros "lizard", is a nodosaurid ankylosaur from the Early to Late Cretaceous period. It is the only known dinosaur species named from terrestrial deposits in Kansas and was designated the official state land fossil in 2023.
Amtosaurus magnus
Amtosaurus () is a genus of ornithischian dinosaur based on a fragmentary skull collected from the Late Cretaceous (Cenomanian to Turonian) Bayanshiree Formation of Mongolia and originally believed to represent an ankylosaurid. Hadrosaurid affinities have also been suggested. However, according to Parish and Barrett, this specimen is too fragmentary to be reliably classified beyond an indeterminate ornithischian. A second species assigned to the genus, A. archibaldi, has become the basis of a valid ankylosaurid taxon, Bissektipelta.
Urbacodon
Urbacodon (meaning "URBAC tooth") is a genus of troodontid theropod dinosaur. Two species have been described, both of which are known from a partial dentary: U. itemirensis from the Dzharakuduk Formation (Cenomanian) of Uzbekistan and U. norelli from the Iren Dabasu Formation of China. Possible remains are also known from the Bissekty Formation (Turonian) of Uzbekistan.
Taurovenator
Taurovenator, from Latin taurus, meaning "bull", and venator, meaning "hunter", is an extinct genus of large, probable carcharodontosaurid theropod dinosaurs that lived in what is now Argentina during the Cenomanian age of the Late Cretaceous, around 95 to 93.9 million years ago. Initially only known from a single postorbital skull bone described from the Huincul Formation, Taurovenator violantei was named in 2016 by Argentine paleontologist Matias Motta and colleagues. Debate came about over the validity of the species, with some paleontologists arguing that Taurovenator was synonymous with t
Sonorasaurus
Sonorasaurus is a genus of brachiosaurid sauropod dinosaur from the Early to Late Cretaceous (Albian to Cenomanian stages, around 112 to 93 million years ago). Its fossils have been found in southern Arizona, United States. Its name, which means "Sonora lizard", comes from the Sonora River that flows in the Sonoran Desert where its fossils were first found. The type species is S. thompsoni, described by Ratkevich in 1998.
Tameryraptor
Tameryraptor ("thief from the beloved land") is an extinct genus of large carcharodontosaurid dinosaur that lived during the Late Cretaceous (Cenomanian age) in what is now Egypt. It is known from a partial skeleton collected in rock layers from the Bahariya Formation by crews of German paleontologist Ernst Stromer in 1914, comprising an incomplete skull, vertebrae, and several other postcranial elements. Stromer described the specimen in 1931, referring it to the previously named Megalosaurus saharicus on the basis of its tooth anatomy, and placing it in a new genus, Carcharodontosaurus. In 1
Fulgurotherium australe
Fulgurotherium (meaning "Lightning beast") is a dubious genus of ornithischian dinosaur from the Late Cretaceous (Cenomanian) Griman Creek Formation. It lived in what is now Australia.
Baotianmansaurus
Baotianmansaurus is an extinct genus of somphospondylian sauropod. Its fossils were discovered in the Late Cretaceous Gaogou Formation of Henan Province, China. The type and only species is B. henanensis — named after the province in which it was found. ==Discovery and naming== The fossils of Baotianmansaurus were discovered at a locality called Neixiang, near Nanyang City in the Henan Province of China. This locality is part of the Baotianman National Nature Reserve corresponds to the rocks of the Gaogou Formation. Dinosaur body fossils in this formation are rare, but a wide variety of egg fo
Alnashetri
Alnashetri is an extinct genus of alvarezsauroid coelurosaurian theropod dinosaur known from the early Late Cretaceous (Cenomanian stage) of La Buitrera, Río Negro Province, Argentina. It contains one known species, Alnashetri cerropoliciensis. Its remains were discovered in Candeleros Formation.
Nopcsaspondylus
Nopcsaspondylus (meaning "Nopcsa's vertebra", in reference to the original describer) is a dubious genus of rebbachisaurid sauropod dinosaur from the Cenomanian-aged (Upper Cretaceous) Candeleros Formation of Neuquén, Argentina. It is based on a now-lost back vertebra described by Nopcsa in 1902 but not named at the time. The specimen had a small vertebral body and large hollows, now known to be typical of rebbachisaurids.
Moros intrepidus
extinct species of tyrannosaur (Tyrannosauroidea)
Burianosaurus
Burianosaurus is an extinct genus of ornithopod dinosaur that lived during the Cenomanian stage of the Late Cretaceous period in what is now the Czech Republic, being the first validly named dinosaur from that country. The type species, B. augustai, was named in 2017; the genus name honours the Czech palaeoartist Zdeněk Burian, and the species name honours the Czech palaeontologist Josef Augusta. The holotype specimen is a femur discovered in sediments belonging to the Peruc-Korycany Formation in 2003, which was described as possibly belonging to an iguanodont in 2005.
Borealosaurus
Borealosaurus is a genus of titanosaurian sauropod dinosaur from the Late Cretaceous of northern China. The type species is Borealosaurus wimani, which was named in 2004.
Meraxes
Meraxes is a genus of large carcharodontosaurid theropod dinosaur from the Late Cretaceous Huincul Formation of Patagonia, Argentina. The genus contains a single species, Meraxes gigas.
Shuangmiaosaurus
Shuangmiaosaurus is a genus of herbivorous iguanodontian ornithopod dinosaur which lived in China during the Late Cretaceous Period, about 100 million years ago.
Beg tse
genus of neoceratopsian dinosaur (fossil)
Elaltitan
Elaltitan is an extinct genus of large lithostrotian titanosaur sauropod dinosaur known from the Late Cretaceous (mid Cenomanian to Turonian stage) of Chubut Province, southern Argentina. It contains a single species, Elaltitan lilloi.
Cathartesaura
Cathartesaura is a genus of rebbachisaurid sauropod dinosaur from Late Cretaceous strata of the Huincul Formation, at the "La Buitrera" locality, in the Neuquén Basin of Río Negro Province, Argentina. The fossil remains, described by Gallina and Apesteguía in 2005, consist of a partial skeleton including vertebrae and limb bones. These were found at the base of the formation, which dates back to the Cenomanian and Turonian epochs, in mudstone and sandstone levels.
Peloroplites
Peloroplites (meaning "monstrous heavy one") is a monospecific genus of nodosaurid dinosaur from Utah that lived during the Late Cretaceous (Cenomanian to lower Turonian stage, 98.2 to 93 Ma) in what is now the Mussentuchit Member of the Cedar Mountain Formation. The type and only species, Peloroplites cedrimontanus, is known from a partial skull and postcranial skeleton. It was named in 2008 by Kenneth Carpenter and colleagues. Peloroplites was 6 metres (20 feet) long and weighed 2 tonnes (4,410 lbs), making it one of the largest known nodosaurids, and came from a time when ankylosaurids and