Eupelycosauria

Eupelycosauria is a large clade of animals characterized by the unique shape of their skull, encompassing all mammals and their closest extinct relatives. They first appeared 308million years ago during the Early Pennsylvanian epoch, with the fossils of Echinerpeton and perhaps an even earlier genus, Protoclepsydrops, representing just one of the many stages in the evolution of mammals, in contrast to their earlier amniote ancestors.

Milosaurus is an extinct genus of non-mammalian synapsids native to Illinois that was alive during the latest Carboniferous and earliest Permian. It was named in 1970 on the basis of FMNH 701, a partial skeleton, as well as referred material.