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Fossil taxa described in 2001

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Masiakasaurus
Masiakasaurus is a genus of small predatory theropod dinosaurs from the Late Cretaceous of Madagascar. In Malagasy, masiaka means "vicious"; thus, the genus name means "vicious lizard". The type species, Masiakasaurus knopfleri, was named after the musician Mark Knopfler, whose music inspired the expedition crew. It was named in 2001 by Scott D. Sampson, Matthew Carrano, and Catherine A. Forster. Unlike most theropods, the front teeth of Masiakasaurus projected forwards instead of straight down. This unique dentition suggests that they had a specialized diet, perhaps including fish and other s
Rapetosaurus
Rapetosaurus ( ) is a genus of titanosaurian sauropod dinosaur that lived in Madagascar from 70 to 66 million years ago, at the end of the Cretaceous Period. Only one species, Rapetosaurus krausei, has been identified. Like other sauropods, Rapetosaurus was a quadrupedal herbivore. It is calculated to have reached lengths of in adulthood.
Paralititan
Paralititan (meaning "tidal giant") is a genus of sauropod dinosaur that lived in present-day Egypt during the Late Cretaceous period. It was described by American paleontologist Joshua B. Smith and colleagues in 2001. The genus contains a single species, Paralititan stromeri, named based on a fragmentary skeleton including vertebrae and limb bones. These fossils were unearthed by an American expedition to the Bahariya Oasis in western Egypt in rock layers of the Bahariya Formation. This formation dates to the Cenomanian stage of the Late Cretaceous, which lasted . An incomplete (back) vertebr
Citipati
Citipati (; meaning "funeral pyre lord") is a genus of oviraptorid dinosaur that lived in Asia during the Late Cretaceous period, about 75 million to 71 million years ago. It is mainly known from the Ukhaa Tolgod locality at the Djadochta Formation, where the first remains were collected during the 1990s. The genus and type species Citipati osmolskae were named and described in 2001. A second species from the adjacent Zamyn Khondt locality may also exist. Citipati is one of the best-known oviraptorids thanks to a number of well-preserved specimens, including individuals found in brooding posit
Ardipithecus kadabba
miocene hominid
Megapnosaurus
Megapnosaurus (meaning "big dead lizard", from Greek μέγα = "big", ἄπνοος = "not breathing", "dead", σαῦρος = "lizard") is an extinct genus of coelophysid theropod dinosaur that lived approximately 188 million years ago during the early part of the Jurassic Period in what is now Africa. The species was a small to medium-sized, lightly built, ground-dwelling, bipedal carnivore, that could grow up to long and weigh up to .
Orrorin
Orrorin is an extinct genus of great ape within the tribe of Hominini from the Miocene Lukeino Formation and Pliocene Mabaget Formation, both of Kenya.
Eotyrannus
Eotyrannus (meaning "dawn tyrant") is a genus of tyrannosauroid theropod dinosaur hailing from the Early Cretaceous Wessex Formation beds, included in Wealden Group, located in the southwest coast of the Isle of Wight, United Kingdom. The remains (MIWG1997.550), consisting of assorted skull, axial skeleton and appendicular skeleton elements, from a juvenile or subadult, found in a plant debris clay bed, were described by Stephen Hutt and colleagues in early 2001. The etymology of the generic name refers to the animal's classification as an early tyrannosaur or "tyrant lizard", while the specif
Hesperosaurus
Hesperosaurus (from Ancient Greek ἕσπερος (hésperos), meaning "western", and σαῦρος (saûros), meaning "lizard") is a herbivorous stegosaurian dinosaur from the Kimmeridgian age of the Jurassic period, approximately 156 million years ago.
Nothronychus
Nothronychus (meaning "slothful claw") is a genus of therizinosaurid theropod dinosaurs that lived in North America during the Late Cretaceous period. The type species, Nothronychus mckinleyi, was described by James Kirkland and Douglas G. Wolfe in 2001. It was recovered near New Mexico's border with Arizona, in an area known as the Zuni Basin, from rocks assigned to the Moreno Hill Formation, dating to the Turonian stage of the Late Cretaceous. A second specimen, described in 2009 as a second species, Nothronychus graffami, was found in the Tropic Shale of Utah, dating to between one million
Vagaceratops
Vagaceratops (meaning "wandering (vagus, Latin) horned face", in reference to its close relationship with Kosmoceratops from Utah) is a genus of chasmosaurine ceratopsid dinosaur which lived during the Late Cretaceous period (late Campanian) in what is now Alberta. Its fossils have been recovered from the Upper Dinosaur Park Formation. It is sometimes included in the genus Chasmosaurus as Chasmosaurus irvinensis instead of being recognized as its own genus.
Aletopelta
Aletopelta (; meaning 'wanderer shield') is a monospecific genus of basal ankylosaurid dinosaur from Southern California that lived during the Late Cretaceous (upper Campanian stage, 75.5 Ma) in what is now the Point Loma Formation. The type and only species, Aletopelta coombsi, is known from a partial skeleton preserving osteoderms. It was originally described in 1996 by W. P. Coombs, Jr. and T.A. Deméré before being named in 2001 by Tracy Ford and James Kirkland. Aletopelta has an estimated size of 5 metres (16 feet) and weight of 2 tonnes (4,409 lbs). The holotype formed a miniature reef an
Khaan
Khaan (; from Mongol 'lord') is an extinct genus of oviraptorid dinosaur known from remains found in the Djadochta Formation of Mongolia, dating to the Late Cretaceous (Campanian, 75-71 million years ago).
Gobisaurus
Gobisaurus is an extinct genus of herbivorous basal ankylosaurid ankylosaur from the Lower Cretaceous of China (Nei Mongol Zizhiqu). The genus is monotypic, containing only the species Gobisaurus domoculus.
Draconyx
Draconyx (meaning "dragon claw") is a genus of ornithopod dinosaur from the Late Jurassic of what is now Portugal. It was found in the Lourinhã Formation in 1991, and described by Octávio Mateus and Miguel Telles Antunes in 2001.
Istiodactylus
Istiodactylus is a genus of pterosaur that lived during the Early Cretaceous period, about 120 million years ago. The first fossil was discovered on the English Isle of Wight in 1887, and in 1901 became the holotype specimen of a new species, O. latidens (Latin for "wide tooth"), in the genus Ornithodesmus. This species was moved to its own genus, Istiodactylus, in 2001; this name is Greek for "sail finger". More specimens were described in 1913, and Istiodactylus was the only pterosaur known from three-dimensionally preserved fossils for much of the 20th century. In 2006, a species from China
Kenyanthropus
Kenyanthropus ('man from Kenya') is a genus of extinct hominin identified from the Lomekwi site by Lake Turkana, Kenya, dated to 3.3 to 3.2 million years ago during the Middle Pliocene. It contains one species, K. platyops, but may also include the two-million-year-old Homo rudolfensis, or K. rudolfensis. Before its naming in 2001, Australopithecus afarensis was widely regarded as the only australopithecine to exist during the Middle Pliocene, but Kenyanthropus evinces a greater diversity than once acknowledged. Kenyanthropus is most recognisable by an unusually flat face and small teeth for s
Losillasaurus
Losillasaurus (meaning "Losilla lizard") is a genus of turiasaurian sauropod dinosaur from the Late Jurassic and possibly Early Cretaceous (Kimmeridgian-?Berriasian) of southeastern Spain. The type species, Losillasaurus giganteus, was discovered in the Villar del Arzobispo Formation in Valencia, and formally described by Casanovas, Santafé, and Sanz in 2001. The holotype material is from a subadult and includes part of a skull; complete cervical, dorsal, sacral, and caudal vertebrae as well as several fragments; skeletal elements from the limbs including a humerus, ulna, radius, and metacarpa
Yanornis martini
Yanornis () is an extinct genus of fish-eating Early Cretaceous birds. Two species have been described, both from Liaoning province, China: Yanornis martini, based on several fossils found in the 120-million-year-old Jiufotang Formation at Chaoyang, and Yanornis guozhangi, from the 124-million-year-old Yixian Formation.
Liaoningosaurus
Liaoningosaurus (meaning "Liaoning lizard") is an unusual genus of basal ankylosaurid dinosaur from the Liaoning Province, China that lived during the Early Cretaceous (late Barremian to early Aptian stages, ~125.4 to 118.9 Ma) in what is now the Yixian and Jiufotang Formation. The type and only species, Liaoningosaurus paradoxus, is known from more than 20 specimens, represented by juvenile individuals. It was named in 2001 by Xu, Wang and You.
Cedarpelta
Cedarpelta is an extinct genus of basal ankylosaurid dinosaur from Utah that lived during the Late Cretaceous period (Cenomanian to lower Turonian stage, 98.2 to 93 Ma) in what is now the Mussentuchit Member of the Cedar Mountain Formation. The type and only species, Cedarpelta bilbeyhallorum, is known from multiple specimens including partial skulls and postcranial material. It was named in 2001 by Kenneth Carpenter, James Kirkland, Don Burge, and John Bird. Cedarpelta has an estimated length of 7 metres (23 feet) and weight of 5 tonnes (11,023 lbs). The skull of Cedarpelta lacks extensi
Eshanosaurus
Eshanosaurus is a genus of a dinosaur from the early Jurassic Period. It is known only from a fossil partial lower jawbone, found in China. It may be a therizinosaurian, and if so the earliest known coelurosaur.
Hadrocodium
Hadrocodium wui is an extinct mammaliaform that lived during the Sinemurian stage of the Early Jurassic approximately in the Lufeng Formation in what is now the Yunnan province in south-western China (, paleocoordinates ). It is considered as the closest relative of the class Mammalia. left|thumb|Life restoration The fossil of this mouse-like, paper-clip sized animal was discovered in 1985 but was then interpreted as a juvenile morganucodontid. Hadrocodium remained undescribed until 2001; since then its large brain and advanced ear structure have greatly influenced the interpretation of the ea
Wellnhoferia
Wellnhoferia (named after Peter Wellnhofer) is a genus of early prehistoric bird-like theropod dinosaur closely related to Archaeopteryx. It is known from a single species, W. grandis, that lived in what is now Germany, during the Late Jurassic. While Wellnhoferia was similar to Archaeopteryx, it had a shorter tail and its fourth toe was shorter than that of Archaeopteryx. Andrzej Elżanowski (2001) of the Institute of Zoology of the University of Wrocław, Poland, determined the differences resulted from a "phylogenetic reduction rather than individual variation."
Jinzhousaurus yangi
Jinzhousaurus is a genus of hadrosauroid dinosaur of the Early Cretaceous of China. The type species is Jinzhousaurus yangi. The generic name refers to the town Jinzhou. The specific name honours Yang Zhongjian is the founder of Chinese paleontology. It was first described by Wang Xiao-lin and Xu Xing in 2001.
Hauffiosaurus
Hauffiosaurus is an extinct genus of Early Jurassic (early Toarcian stage) pliosaurid plesiosaur known from Holzmaden of Germany and from Yorkshire of the United Kingdom. It was first named by Frank Robin O’Keefe in 2001 and the type species is Hauffiosaurus zanoni. In 2011, two additional species were assigned to this genus: H. longirostris and H. tomistomimus.
Jiangshanosaurus
Jiangshanosaurus is a genus of herbivorous titanosauriform sauropod dinosaur that lived in China approximately 92-88 million years ago, during the Turonian and Coniacian stages of the Late Cretaceous.
Venenosaurus
Venenosaurus ( ) is a genus of sauropod dinosaur that lived in what is now Utah during the Early Cretaceous. Its type and only species is Venenosaurus dicrocei. Fossils of Venenosaurus were first discovered in 1998, by Denver Museum of Natural History volunteer Anthony DiCroce, and described as a new genus and species in 2001 by Virginia Tidwell and colleagues, who named the species for DiCroce. Venenosaurus was a relatively small sauropod, and was similar to Cedarosaurus, another sauropod from the Early Cretaceous of Utah.
Pezosiren
Pezosiren portelli, also known as the "walking manatee", is a basal sirenian from the early Eocene of Jamaica, 50 million years ago. The type specimen is represented by a Jamaican fossil skeleton, described in 2001 by Daryl Domning, a marine mammal paleontologist at Howard University in Washington, DC. It is believed to have had a hippopotamus-like amphibious lifestyle, and is considered a transitional form between land and sea mammals.
Neimongosaurus
Neimongosaurus (meaning "Nei Mongol lizard") is a genus of herbivorous therizinosaurian theropod dinosaur that lived in China during the Late Cretaceous period. Its fossils are known from the strata of the Iren Dabasu Formation. It is known from two specimens, discovered in 1999 by researchers from the Ministry of Land and Resources and described two years later. One species, N. yangi, is known, named after Chinese palaeontologist Yang Zhongjian.
Quilmesaurus
Quilmesaurus is a genus of carnivorous theropod dinosaur from the Upper Cretaceous (Campanian to Maastrichtian stages) Allen Formation of Argentina. It was a member of Abelisauridae, closely related to genera such as Carnotaurus. The only known remains of this genus are leg bones which share certain similarities to a variety of abelisaurids. However, these bones lack unique features, which may render Quilmesaurus a nomen vanum (more commonly known as a nomen dubium, or "dubious name").
Pukyongosaurus
Pukyongosaurus (meaning "Pukyong lizard", after the Pukyong National University) is a genus of titanosauriform dinosaur that lived in South Korea during the Early Cretaceous Period (Aptian - Albian). It may have been closely related to Euhelopus, and is known from a series of vertebrae in the neck and back. The characteristics that were originally used to distinguish this genus have been criticized as being either widespread or too poorly preserved to evaluate, rendering the genus an indeterminate nomen dubium among titanosauriforms. A 2022 study noted that Pukyongosaurus is probably a somphos
Tyrannasorus rex
extinct species of beetle
Viti Levu Giant Pigeon
species of bird (fossil)
Ocepeia
Ocepeia is an extinct genus of afrotherian mammal that lived in present-day Morocco during the middle Paleocene epoch, approximately 60 million years ago. First named and described in 2001, the type species is O. daouiensis from the Selandian stage of Morocco's Ouled Abdoun Basin. A second, larger species, O. grandis, is known from the Thanetian, a slightly younger stage in the same area. In life, the two species are estimated to have weighed about and , respectively, and are believed to have been specialized leaf-eaters. The fossil skulls of Ocepeia are the oldest known afrotherian
Planicoxa venenica
Planicoxa is a genus of advanced iguanodontian dinosaur from the Early Cretaceous of North America. It is known from the partial skeletons of several individual specimens. Its fossils were discovered in Utah, United States.
Akmonistion
Akmonistion is an extinct genus of cartilaginous fish that lived in the Early Carboniferous. The genus contains a single species, A. zangerli, discovered by Stan Wood in 1982. It is distinguished by an unusual enlarged formation of the dorsal fin, called a "spine-brush complex", of unknown function. This is also found in the better known genus Stethacanthus. Remains have only been found near Bearsden in Scotland, where a complete fossil was uncovered in 1982. It reached in length. The genus name is derived from the Ancient Greek “akmon” (“anvil”) + “istion” (“sail”) referring to the appearance
Bienosaurus
Bienosaurus (meaning "Boien's lizard) is a dubious genus of thyreophoran dinosaur from the Early Jurassic (probably Hettangian to Sinemurian) Lower Lufeng Formation of Yunnan Province in China.
Longipteryx
Longipteryx is a genus of prehistoric bird which lived during the Early Cretaceous (Aptian stage, 120.3 million years ago). It contains a single species, Longipteryx chaoyangensis. Its remains have been recovered from the Jiufotang Formation at Chaoyang in Liaoning Province, China. Apart from the holotype IVPP V 12325 - a fine and nearly complete skeleton — another entire skeleton (IPPV V 12552), some isolated bones (a humerus and furcula, specimens IPPV V 12553, and an ulna, IPPV V 12554) and many other specimens are known to date.
Haopterus
Haopterus is a genus of pterodactyloid pterosaur from the Barremian-Aptian-age Lower Cretaceous Yixian Formation of Liaoning, China. Its fossil remains dated back 124.6 million years ago.
Hypuronector
Hypuronector is a genus of extinct drepanosaur reptile from the Late Triassic Lockatong Formation of New Jersey. The etymology of the name translates as "deep-tailed swimmer from the lake", in reference to its assumed aquatic habits hypothesized by its discoverers. Hypuronector was related to the arboreal Megalancosaurus. It was a small animal, estimated to be only long in life. So far dozens of specimens of Hypuronector are known, though scientists have not found any complete skeletons. This makes attempts to reconstruct Hypuronector's body or lifestyle highly speculative and controversial.
Ruehleia
Ruehleia is a genus of sauropodomorph dinosaur from the Late Triassic period of Germany. The type species is R. bedheimensis, described by Galton in 2001, and is named for the German paleontologist Hugo Rühle von Lilienstern. Discovered in 1952, the holotype consists of one nearly complete skeleton, including cervical (neck), dorsal (back), and caudal (tail) vertebrae; a partial sacrum; a scapulocoracoid; pelvic bones; most limb bones; and a partially complete manus (hands). The fossils were found in the Trossingen Formation of central Germany and date to the Norian stage, around 216 to 208 mi
Stratiotosuchus
Stratiotosuchus (from Greek, (stratiōtēs, "soldier") and (soûchos, "crocodile")) is an extinct genus of baurusuchid mesoeucrocodylian from the Adamantina Formation in Brazil. It lived during the Late Cretaceous. The first fossils were found in the 1980s, and the type species Stratiotosuchus maxhechti was named in 2001. A hyperpredator, it and other baurusuchids may have filled niches occupied elsewhere by theropod dinosaurs.
Rubricacaecilia
Rubricacaecilia ("Red Earth Caecilian") is an extinct genus of caecilian (limbless amphibian) from the Berriasian aged Ksar Metlili Formation in Morocco. The genus contains a single species, Rubricacaecilia monbaroni, Named for geologist M. monbaron. It is known from a nearly complete pseudodentary bone, several fragmentary pseudodentaries, a partial pseudangular, a partial palatine, an atlas, and vertebrae.
Apsaravis ukhaana
Apsaravis is a Mesozoic avialan genus from the Late Cretaceous. The single known species, Apsaravis ukhaana, lived about 78 million years ago, in the Campanian age of the Cretaceous period. Its fossilized remains were found in the Camel's Humps sublocality of the Djadokhta Formation, at Ukhaa Tolgod in the Gobi Desert of Mongolia. They were collected in the 1998 field season by the Mongolian Academy of Sciences/American Museum of Natural History Paleontological Expeditions. It was described by Norell and Clarke (2001).
Prozostrodon brasiliensis
Prozostrodon is an extinct genus of probainognathian cynodonts that was closely related to mammals. The remains were found in Brazil and are dated to the Carnian age of the Late Triassic. The holotype has an estimated skull length of , indicating that the whole animal may have been the size of a cat. The teeth were typical of advanced cynodonts, and the animal was probably a carnivore hunting reptiles and other small prey.
Abdounodus
Abdounodus ("Abdoun tooth") is an extinct genus of mammal known from the middle Paleocene of Northern Africa. The sole species, A. hamdii, is known from teeth and jaw bones discovered in the Ouled Abdoun Basin of present-day Morocco in 2001.
Kalimantsia bulgarica
Kalimantsia is an extinct chalicothere from the Miocene of Bulgaria, Europe. It contains one species, Kalimantsia bulgarica.
Bageherpeton
Bageherpeton is an extinct genus of archegosaurid temnospondyl amphibian from the Upper Permian period of Rio Grande do Sul, Brazil (Paleorrota). It was found in the Rio do Rasto Formation, and was named in honor of the city of Bagé near where it was found. It was described from a lower jaw.
Pabwehshi
Pabwehshi (meaning "Pab [Formation] beast ["wehshi" in Urdu]") is an extinct genus of baurusuchine mesoeucrocodylian. It is based on GSP-UM 2000, a partial snout and corresponding lower jaw elements, with another snout assigned to it. These specimens were found in Maastrichtian-age Upper Cretaceous rocks of the Vitakri and Pab formations in Balochistan, Pakistan, and represent the first diagnostic crocodyliform fossils from Cretaceous rocks of South Asia. Pabwehshi had serrated interlocking teeth in its snout that formed a "zig-zag" cutting edge. Pabwehshi was named in 2001 by Jeffrey A.
Chrysocetus
Chrysocetus is a genus of extinct archaeocete ceteceans belonging to the Basilosauridae that is known from the Late Eocene of the eastern United States and western Africa. Across its range two species are known, the American Chrysocetus healyorum and the African Chrysocetus fouadassii, named in 2001 and 2015 respectively. Chrysocetus was a comparably small basilosaurid, larger than Saghacetus and Tutcetus but smaller than most other members of this group. In addition to being noted for its well-preserved hip bones, Chrysocetus is unique amongst basilosaurids for possibly having only had a sing
Artiocetus
Artiocetus is an extinct genus of early whales belonging to the family Protocetidae. It was a close relative to Rodhocetus and its tarsals indicate it resembled an artiodactyl.
Yuyuanozoon
Yuyuanozoon magnificissimi, from the Cambrian Stage 3 Chengjiang lagerstatte, is the largest known vetulicolian, an extinct species of marine animal, with specimens up to 20 cm in length compared to 5–14 cm for other vetulicolian species.
Yixianornis
Yixianornis (meaning "Yixian Formation bird") is a bird genus from the early Cretaceous period. Its remains have been found in the Jiufotang Formation at Chaoyang (People's Republic of China) dated to the early Aptian age, around 120 million years ago. Only one species, Yixianornis grabaui, is known at present. The specific name, grabaui, is named after American paleontologist Amadeus William Grabau, who surveyed China in the early 20th century.
Lumkuia
Lumkuia is an extinct genus of cynodont, fossils of which have been found in the Cynognathus Assemblage Zone of the Beaufort Group in the South African Karoo Basin that date back to the early Middle Triassic. It contains a single species, Lumkuia fuzzi, which was named in 2001 on the basis of the holotype specimen BP/1/2669, which can now be found at the Bernard Price Institute in Johannesburg, South Africa. The genus has been placed in its own family, Lumkuiidae. Lumkuia is not as common as other cynodonts from the same locality such as Diademodon and Trirachodon.
Riograndia
Riograndia is an extinct genus of prozostrodontian cynodonts from the Late Triassic of Brazil. The type and only species is Riograndia guaibensis, named after the State of Rio Grande do Sul and Guaíba Basin, where it was discovered. Remains have been found in the Caturrita Formation of the geopark of Paleorrota. It was a small non-mammalian cynodont, with several advanced features also present in mammals. Several specimens of Riograndia guaibensis have been found in the towns of Candelária and Faxinal do Soturno in the Caturrita Formation. The genus defines the Riograndia Assemblage Zone.
Cherninia
Cherninia is an extinct genus of mastodonsaurid temnospondyl known from the Denwa Formation of India and the Ntawere Formation of Zambia.
Cearachelys
Cearachelys is an extinct genus of pleurodiran turtle which existed some 110 million years ago. The genus is monotypic, with only type species Cearachelys placidoi known.
Iberospondylus
Iberospondylus is an extinct genus of basal temnospondyl amphibian which lived in a marine environment. The type material was found in the Emma Quarry Amphibian Bed of the Puertollano Basin, Ciudad Real province, southern Spain and extended the record for temnospondyls on the peninsula by 45 million years. Along with the holotype, a skull with several disarticulated vertebrae and ribs, two other partial skeletons are known. The name is derived from "Iberia" name of the peninsula where Spain is located, plus "spodylos", Greek for vertebra. The species name is in honor of Dr. Hans-Peter Schultze
Limenavis
Limenavis is a genus of ornithuran dinosaurs from the Late Cretaceous. It lived about 70 million years ago, around the Campanian-Maastrichtian boundary. Known from several broken bones, the remains of the only known species Limenavis patagonica were found in rocks of the "lower member" of the Allen Formation at Salitral Moreno, 20 km south of General Roca, Río Negro (Argentina). It is one of the closest relatives, in the fossil record, of the modern birds.