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Prehistoric pseudosuchian genera

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Deinosuchus
Deinosuchus is an extinct genus of eusuchian, either an alligatoroid crocodilian or a stem-group crocodilian, which lived during the Late Cretaceous around . The first remains were discovered in North Carolina (United States) in the 1850s, and the genus was first described in 1909. Additional fragments were discovered in the 1940s and were later incorporated into an influential, though inaccurate, skull reconstruction at the American Museum of Natural History. Knowledge of Deinosuchus remains incomplete, but better cranial material found in recent years has expanded scientific understanding of
Sarcosuchus
Sarcosuchus (), from Ancient Greek σάρξ (sárx), meaning "flesh", and Σοῦχος (Soûkhos), meaning "Sobek", is an extinct genus of crocodyliform that lived during the Early Cretaceous, from the late Hauterivian to the early Albian stages, 130 to 112 million years ago of what is now Africa and South America. It was one of the largest pseudosuchians, with the largest specimen of S. imperator reaching approximately long and weighing up to . It is known from two species; S. imperator from the early Albian Elrhaz Formation of Niger, and S. hartti from the Late Hauterivian of northeastern Brazil. Other
Postosuchus
Postosuchus, meaning "Crocodile from Post", is an extinct genus of rauisuchid reptiles comprising two species, P. kirkpatricki and P. alisonae, that lived in what is now North America during the Late Triassic. Postosuchus is a member of the clade Pseudosuchia, the lineage of archosaurs that includes modern crocodilians (the other main group of archosaurs is Avemetatarsalia, the lineage that includes all archosaurs more closely related to birds than to crocodilians). Its name refers to Post Quarry, a place in Texas where many fossils of the type species, P. kirkpatricki, were found.
Desmatosuchus
Desmatosuchus (, from Greek δεσμός desmos 'link' + σοῦχος soûkhos 'crocodile') is an extinct genus of archosaur belonging to the Order Aetosauria. It lived in North America and potentially India (cf. Desmatosuchus) during the Late Triassic.
Purussaurus
Purussaurus is a genus of extinct giant caimans that lived in the Americas during the Miocene epoch, from the Friasian to the Huayquerian in the SALMA classification. It is known from skull material found in the Brazilian and Peruvian Amazon, Argentina, Colombian Villavieja Formation, Panamanian Culebra Formation, Urumaco, and Socorro Formations of northern Venezuela.
Dakosaurus
Dakosaurus is an extinct genus of crocodylomorph within the family Metriorhynchidae that lived during the Late Jurassic and Early Cretaceous. It was large, with teeth that were serrated and compressed lateromedially (flattened from side to side). The genus was established by Friedrich August von Quenstedt in 1856 for an isolated tooth named Geosaurus maximus by Theodor Plieninger in 1846. Dakosaurus was a carnivore that spent much, if not all, of its life out at sea. The extent of its adaptation to a marine lifestyle means that it is most likely that it mated at sea, but since no eggs or nests
Ornithosuchus
Ornithosuchus (from , "bird" and , "crocodile") is an extinct genus of pseudosuchians from the Late Triassic (Carnian) Lossiemouth Sandstone of Scotland. It was originally thought to be the ancestor to the carnosaurian dinosaurs (such as Allosaurus), but it is now known to be more closely related to crocodilians than to dinosaurs.
Rhamphosuchus
Rhamphosuchus is a genus of extinct gavialoid crocodylians from the Indian subcontinent. Two species are currently recognized, the geologically older R. pachyrhynchus, represented by fossils recovered in Oligocene and Miocene deposits of Pakistan's Bugti and Laki Hills, and the younger R. crassidens, known from the Pliocene Siwalik Hills of Northern India. Additional remains are also known from the Pliocene of Nepal and a tentative third species was recovered in Pakistan. Both species are known for their great size, with rigorous estimates showing potential body lengths ranging from for R. cra
Saurosuchus
Saurosuchus (meaning "lizard crocodile") is an extinct genus of large loricatan pseudosuchian archosaurs that lived in South America during the Late Triassic period. The first and holotype specimen was discovered in 1957 by Galileo J. Scaglia and Leocadio Soria in the strata of the Ischigualasto Formation in northwestern Argentina. The type and only species, S. galilei, was named in 1959 by Osvaldo A. Reig. Several specimens, including an adult and multiple subadults, have been discovered since.
Araripesuchus
Araripesuchus is a genus of extinct crocodyliform that existed during the Cretaceous period of the late Mesozoic era some 125 to 66 million years ago. Araripesuchus is generally considered to be a notosuchian (belonging to the clade Mesoeucrocodylia), characterized by the varied teeth types and distinct skull elements. Seven species have been referred to Araripesuchus, though it has been argued that the phylogenetic position of this genus is uncertain, and that taxonomic revision is required.
Simosuchus
Simosuchus is an extinct genus of notosuchian crocodyliforms from the Late Cretaceous of Madagascar. It is named for its unusually short skull. Fully grown individuals were about in length. The type species is Simosuchus clarki, found from the Maevarano Formation in Mahajanga Province, although one isolated multicuspid tooth of this genus was discovered in Kallamedu Formation of India.
Machimosaurus
Machimosaurus is an extinct genus of machimosaurid crocodyliform from the Late Jurassic (Kimmeridgian and Tithonian) and Early Cretaceous. The type species, Machimosaurus hugii, was found in Switzerland. Other fossils have been found in England, France, Germany, Portugal, Switzerland and Tunisia. Machimosaurus rex is the largest named teleosauroid and thalattosuchian, with an estimated length of up to (skull length ). Machimosaurus is the largest known crocodyliform of the Jurassic.
Kaprosuchus
Kaprosuchus is an extinct genus of mahajangasuchid crocodyliform. It is known from a single nearly complete skull collected from the Upper Cretaceous Echkar Formation of Niger. The name means "boar crocodile" from the Greek , kapros ("boar") and , soukhos ("crocodile") in reference to its unusually large caniniform teeth which resemble those of a boar. It has been nicknamed "BoarCroc" by Paul Sereno and Hans Larsson, who first described the genus in a monograph published in ZooKeys in 2009 along with other Saharan crocodyliforms such as Anatosuchus and Laganosuchus. The type species is K. saha
Goniopholis
Goniopholis (meaning "angled scale") is an extinct genus of goniopholidid crocodyliform that lived in Europe and North America during the Late Jurassic and Early Cretaceous. Like other goniopholidids, it resembled living crocodilians, and probably had a similar ecology as semi-aquatic ambush predators.
Poposaurus
Poposaurus is an extinct genus of pseudosuchian archosaur from the Late Triassic of the southwestern and eastern United States. It belongs to the clade Poposauroidea, an unusual group of Triassic pseudosuchians that includes sail-backed, beaked, and aquatic forms. Fossils have been found in Wyoming, Utah, Arizona, Texas, and Virginia. Except for the skull, most parts of the skeleton are known. The type species, P. gracilis, was described and named by Maurice Goldsmith Mehl in 1915. A second species, P. langstoni, was originally the type species of the genus Lythrosuchus. Since it was first des
Batrachotomus
Batrachotomus is a genus of prehistoric archosaur. Fossils of this animal have been found in southern Germany and dated from the Ladinian stage of the Middle Triassic period, around 242 to 237 million years ago. Batrachotomus was described by palaeontologist David J. Gower 22 years after its discovery.
Teratosaurus
Teratosaurus is a genus of rauisuchians known from the late Triassic Stubensandstein (Löwenstein Formation, Norian stage) of Germany. It is estimated to be in length.
Arizonasaurus
Arizonasaurus (meaning "Arizona reptile") is a genus of ctenosauriscid archosaur from the Middle Triassic (243 million years ago) Moenkopi Formation of what is now Arizona. It is known for having a large back sail formed by elongated neural spines of its vertebrae. The type and only known species is A. babbitti, named and described in 1947.
Geosaurus
Geosaurus is an extinct genus of marine crocodyliform within the family Metriorhynchidae, that lived during the Late Jurassic and the Early Cretaceous. Geosaurus was a carnivore that spent much, if not all, its life out at sea. No Geosaurus eggs or nests have been discovered, so little is known of the reptile's lifecycle, unlike other large marine reptiles of the Mesozoic, such as plesiosaurs or ichthyosaurs which are known to give birth to live young out at sea. Where Geosaurus mated, whether on land or at sea, is currently unknown. The name Geosaurus means "Mother of Giants lizard", and is d
Metriorhynchus
Metriorhynchus is an extinct genus of marine crocodyliform that lived in the oceans during the Late Jurassic. The type species, M. brevirostris was named in 1829 as a species of Steneosaurus before being named as a separate genus by the German palaeontologist Christian von Meyer in 1832. The name Metriorhynchus means "moderate snout", and is derived from the Greek Metrio- ("moderate") and -rhynchos ("snout").
Quinkana
Quinkana is an extinct genus of mekosuchine crocodylians that lived in Australia from about 25 million to about 10,000 years ago, with the majority of fossils having been found in Queensland. Four species are currently recognized, all of which have been named between 1981 and 1997. The two best understood species are Q. fortirostrum, the type species, and Q. timara, a more gracile form from the Miocene. The other two species, Q. babarra and Q. meboldi, from the Pliocene and Oligocene respectively, are only known from a few poorly preserved bone fragments. The name Quinkana comes from the "Quin
Pristichampsus
Pristichampsus (from , 'saw' and , 'crocodile') is a non-diagnostic and potentially dubious extinct genus of crocodylian from France and possibly also Kazakhstan that is part of the monotypic Pristichampsidae family. As the type species, Pristichampsus rollinatii, was based on insufficient material when described in 1831 and 1853, the taxonomic status of the genus is in doubt, and other species have been referred to other genera, primarily Boverisuchus.
Armadillosuchus arrudai
Armadillosuchus is an extinct genus of sphagesaurid crocodylomorph. It was described in February 2009 from the late Campanian to early Maastrichtian Adamantina Formation of the Bauru Basin in Brazil, dating to approximately 70 Ma. Armadillosuchus was among the larger and more robust sphagesaurids, with a total length of approximately .
Allodaposuchus
Allodaposuchus is an extinct genus of crocodyliforms that lived in what is now southern Europe during the Campanian and Maastrichtian stages, and possibly the Santonian stage, of the Late Cretaceous. Although generally classified as a non-crocodylian eusuchian crocodylomorph, it is sometimes placed as one of the earliest true crocodylians. Allodaposuchus is one of the most common Late Cretaceous crocodylomorphs from Europe, with fossils known from Romania, Spain, and France.
Stomatosuchus inermis
Stomatosuchus (meaning "mouth crocodile") is an extinct stomatosuchid neosuchian from the Late Cretaceous (Cenomanian) Bahariya Formation of Egypt. The type and only species is S. inermis. Much of what is known about Stomatosuchus has been inferred from the related genus Laganosuchus.
Borealosuchus
Borealosuchus (meaning "northern crocodile") is an extinct genus of crocodyliforms that lived from the Late Cretaceous to the Eocene in North America. It was named by Christopher Brochu in 1997 for several species that had been assigned to Leidyosuchus. The species assigned to it are: B. sternbergii, the type species, from the Maastrichtian (Late Cretaceous) of Colorado, Montana, North Dakota, South Dakota, and Wyoming; B. acutidentatus, from the Paleocene of Saskatchewan; B. formidabilis, from the Paleocene of North Dakota; B. griffithi, from the Paleocene of Alberta; and B. wilsoni, from the
Aetosaurus
Aetosaurus is an extinct genus of pseudosuchian reptile belonging to the order Aetosauria. It is generally considered to be the most primitive aetosaur. Three species are currently recognized: A. ferratus, the type species from Germany and Italy; A. crassicauda from Germany; and A. arcuatus from eastern North America. Additional specimens referred to Aetosaurus have been found in the Chinle Group of the southwestern United States, and the Fleming Fjord Formation of Greenland. Specimens of Aetosaurus occur in Norian-age strata.
Stagonolepis
Stagonolepis is an extinct genus of stagonolepidid aetosaur known from the Late Triassic (Carnian stage) Hassberge Formation of Germany, the Drawno Beds of Poland, and the Lossiemouth Sandstone of Scotland. Supposed fossils from North and South America have been placed into their own genera, Calyptosuchus and Aetosauroides, respectively.
Gryposuchus
Gryposuchus (meaning "hooked crocodile") is an extinct genus of gavialid crocodilian. Fossils have been found from Argentina, Colombia, Venezuela, Brazil and the Peruvian Amazon. The genus existed during the Miocene epoch (Colhuehuapian to Huayquerian). One recently described species, G. croizati, grew to an estimated length of . Gryposuchus is the type genus of the subfamily Gryposuchinae, although a 2018 study indicates that Gryposuchinae and Gryposuchus might be paraphyletic and rather an evolutionary grade towards the gharial.
Protosuchus
Protosuchus (from , "first" and , "crocodile") is an extinct genus of carnivorous crocodyliform from the Early Jurassic. It is among the earliest animals that resemble crocodilians. Protosuchus was about in length thumb|left|The skull of Protosuchus richardsoni (AMNH 3024) thumb|left|The pelvis and hindlimbs of Protosuchus richardsoni (AMNH 3024) thumb|right|Protosuchus richardsoni fossil AMNH 3024 As an early crocodilian relative, its skull and postcranial anatomy featured more crocodilian characteristics than its earlier ancestors; it had short jaws that broadened out at the base of the skul
Baurusuchus
left|thumb|450x450px|Skeletal diagram and size comparison showing some of the different Baurusuchus specimens in scientific literature (human: 175 cm tall). Several more specimens have been discovered and stored in museums Baurusuchus is an extinct genus of baurusuchid mesoeucrocodylian, which lived in Brazil from 90 to 83.5 million years ago, in the Late Cretaceous period. It was a terrestrial predator, estimated to reach up to in weight. Baurusuchus lived during the Turonian to Santonian stages of the Late Cretaceous Period, in Adamantina Formation, Brazil. It gets its name from the Brazilia
Hesperosuchus
Hesperosuchus is an extinct genus of crocodylomorph reptile that contains a single species, Hesperosuchus agilis. Remains of this pseudosuchian have been found in Late Triassic (Carnian) strata from Arizona and New Mexico. Because of similarities in skull and neck anatomy and the presence of hollow bones, Hesperosuchus was formerly thought to be an ancestor of later carnosaurian dinosaurs, but based on more recent findings and research it is now known to be more closely related to crocodilians rather than dinosaurs.
Anatosuchus
Anatosuchus ("duck crocodile", the name from the Latin anas ("duck") and the Greek souchos ("crocodile"), for the broad, duck-like snout) is an extinct genus of notosuchian crocodyliforms discovered in Gadoufaoua, Niger, and described by a team of palaeontologists led by the American Paul Sereno in 2003, in the Journal of Vertebrate Paleontology.
Prestosuchus
Prestosuchus (meaning "Prestes crocodile") is an extinct genus of pseudosuchian in the group Loricata, which also includes Saurosuchus and Postosuchus. It has historically been referred to as a "rauisuchian", and was the defining member of the family Prestosuchidae, though the validity of both of these groups is questionable: Rauisuchia is now considered paraphyletic and Prestosuchidae is polyphyletic in its widest form.
Shuvosaurus
Shuvosaurus (meaning "Shuvo [Chatterjee]'s lizard") is a genus of beaked, bipedal poposauroid pseudosuchian from the Late Triassic (early to middle Norian) of western Texas. Despite superficially resembling a theropod dinosaur, especially the ostrich-like ornithomimids, it is instead more closely related to living crocodilians than to dinosaurs. Shuvosaurus is known by the type and only species S. inexpectatus, and is closely related to the very similar Effigia within the clade Shuvosauridae. Shuvosaurus was originally described from a restored skull and very few fragmentary postcranial bones
Laganosuchus
Laganosuchus is an extinct genus of stomatosuchid crocodyliform. Fossils have been found from Niger and Morocco and date back to the Upper Cretaceous.
Mahajangasuchus
Mahajangasuchus is an extinct genus of crocodyliform which had blunt, laterally compressed and serrated teeth. The type species, M. insignis, lived during the Late Cretaceous; its fossils have been found in the Maevarano Formation in northern Madagascar. It was a fairly large predator, measuring up to long.
Teleosaurus
Teleosaurus (from , 'perfect' and , 'lizard') is an extinct genus of teleosaurid crocodyliform found in the Middle Jurassic Calcaire de Caen Formation of France. It was approximately in length. The holotype is MNHN AC 8746, a quarter of a skull and other associated postcranial remains, while other fragmentary specimens are known. The type species is T. cadomensis, but a second species, T. geoffroyi may also exist. It was previously considered a wastebasket taxon, with many other remains assigned to the genus.
Effigia okeeffeae
Effigia is an extinct genus of shuvosaurid known from the Late Triassic of New Mexico, south-western USA. With a bipedal stance, long neck, and a toothless beaked skull, Effigia and other shuvosaurids bore a resemblance to the ornithomimid dinosaurs of the Cretaceous Period. However, shuvosaurids were not dinosaurs, but were instead a specialized family of poposauroid pseudosuchians, meaning that their closest living relatives are crocodilians.
Theriosuchus
Theriosuchus is an extinct genus of atoposaurid neosuchian from Late Jurassic to Early Cretaceous of Europe (Hungary & southern England), Southeast Asia (Thailand) and western North America (Wyoming), with fragmentary records from Middle Jurassic and Early Cretaceous sites in China, Morocco, and Scotland.
Saltoposuchus
Saltoposuchus is an extinct genus of small (1–1.5 m and 10–15 kg), long-tailed crocodylomorph reptile (Sphenosuchia), from the Norian (Late Triassic) of Europe. The name translated means "leaping foot crocodile". It has been proposed that Terrestrisuchus gracilis and Saltoposuchus connectens represent different ontogenetic stages of the same genus. Saltoposuchus was commonly (and incorrectly) referred to in popular literature as the ancestor (or close ancestors) to dinosaurs; however, recent scientific research shows that this is not the case.
Ticinosuchus
Ticinosuchus is an extinct genus of suchian archosaur (distantly related to modern crocodilians) from the Middle Triassic (Anisian–Ladinian) of Switzerland and Italy. The genus name means "Ticino crocodile" due to its origin from the Swiss canton Ticino. It is one of the only terrestrial animals found in the Besano Formation, a fossil-rich lagerstätte on Monte San Giorgio renowned for its marine fauna (including different groups of fossil fishes, ichthyosaurs, tanystropheids, nothosaurs and placodonts, among others).
Bernissartia
Bernissartia ('of Bernissart') is an extinct genus of neosuchian crocodyliform that lived in the Early Cretaceous, around 130 million years ago.
Rauisuchus
Rauisuchus (meaning "Wilhelm Rau's crocodile") is a genus of extinct archosaurs which lived in what is now the Geopark of Paleorrota (Santa María Formation), Brazil, during the Late Triassic period (235–228 million years ago). It contains one species, R. tiradentes.F. v. Huene. (1942) Die fossilen Reptilien des südamerikanischen Gondwanalandes. Ergebnisse der Sauriergrabung in Südbrasilien 1928/29. München: C.H. Beck'sche Verlagsbuchhandlung
Fasolasuchus
Fasolasuchus is an extinct genus of loricatan. Fossils have been found in the Los Colorados Formation of the Ischigualasto-Villa Unión Basin in northwestern Argentina that date back to the Norian stage of the Late Triassic, making it one of the last rauisuchians (an informal grouping of large carnivourous archosaurs) to have existed before rauisuchians became extinct at the end of the Triassic.
Gracilisuchus
Gracilisuchus (meaning "slender crocodile") is an extinct genus of tiny pseudosuchian (a group which includes the ancestors of crocodilians) from the Late Triassic of Argentina. It contains a single species, G. stipanicicorum, which is placed in the clade Suchia, close to the ancestry of crocodylomorphs. Both the genus and the species were first described by Alfred Romer in 1972.
Steneosaurus
Steneosaurus (from , 'narrow' and , 'lizard') is a dubious genus of teleosaurid crocodyliform from the Middle or Late Jurassic (Callovian or early Oxfordian) of France and possibly also India. The genus has been used as a wastebasket taxon for thalattosuchian fossils for over two centuries, and almost all known historical species of teleosauroid have been included within it at one point. The genus has remained a wastebasket, with numerous species still included under the label '''Steneosaurus''', many of which are unrelated to each other (either paraphyletic or polyphyletic with respect to eac
Rhamphocephalus
Rhamphocephalus ("beak head") is an extinct genus of fossil reptile from the Middle Jurassic (Bathonian stage) Cotswold Slate Formation of Gloucestershire, England. The name was erected as a genus of pterosaur and became a 'wastebasket taxon' for British Jurassic pterosaur remains until a recent revision. Rhamphocephalus comprises several named species, two of which are pterosaurian, but the type species - R. prestwichi - is based on remains now identified as a thalattosuchian. Because it is poorly preserved and lacks features that distinguish it from other thalattosuchians, R. prestwichi is c
Thoracosaurus
Thoracosaurus (chest lizard) is an extinct genus of long-snouted eusuchian which existed during the Late Cretaceous and Early Paleocene in North America and Europe.
Isisfordia
Isisfordia is an extinct genus of crocodyliform closely related to crocodilians that lived in Australia during the Middle Cretaceous (Albian–Cenomanian).
Notosuchus
Notosuchus (; 'southern crocodile') is an extinct genus of South American notosuchian crocodyliforms. It was terrestrial, living approximately 85 million years ago in the Santonian stage of the Late Cretaceous.
Carnufex
Carnufex is an extinct genus of crocodylomorph suchian from the Late Triassic of North America. The genus was first described in 2015 by Zanno et al., who named the binomial Carnufex carolinensis, meaning "Carolina butcher". Two specimens are known, the holotype skull and skeleton NCSM 21558, and the referred humerus NCSM 21623. The specimens are from the Carnian-age Pekin Formation, which dates to 231 million years ago. Based on the holotype, Carnufex would have been about long and tall, although it may have gotten larger due to the holotype not being fully grown.
Pakasuchus
Pakasuchus is a genus of notosuchian crocodyliform distinguished by its unusual mammal-like appearance, including mammal-like teeth that would have given the animal the ability to chew. It also had long, slender legs and a doglike nose. Fossils have been found in the Galula Formation of Rukwa Rift Basin of southwestern Tanzania, and were described in 2010 in the journal Nature. Pakasuchus is originally considered to lived approximately 105 million years ago, in the mid-Cretaceous, but later age of site is reconsidered to the late Cretaceous, Cenomanian to Campanian instead. The type species is
Pelagosaurus
Pelagosaurus (meaning "lizard of the open sea") is an extinct genus of thalattosuchian crocodyliform that lived during the Toarcian stage of the Lower Jurassic, around 183 Ma to 176 Ma (million years ago), in shallow epicontinental seas that covered much of what is now Western Europe. The systematic taxonomy of Pelagosaurus has been fiercely disputed over the years, and was assigned to Thalattosuchia after its systematics within Teleosauridae were disputed. Pelagosaurus measured long.
Nundasuchus
Nundasuchus is an extinct genus of crurotarsan, possibly a suchian archosaur related to Paracrocodylomorpha. Remains of this genus are known from the Middle Triassic Manda beds of southwestern Tanzania. It contains a single species, Nundasuchus songeaensis, known from a single partially complete skeleton, including vertebrae, limb elements, osteoderms, and skull fragments.
Mekosuchus
Mekosuchus is a genus of extinct Australasian mekosuchine crocodilian. Species of Mekosuchus were generally small-sized (less than long), terrestrial animals with short, blunt-snouted heads and strong limbs. Four species are currently recognized, M. inexpectatus, M. whitehunterensis, M. sanderi and M. kalpokasi, all known primarily from fragmentary remains.
Aegisuchus
Aegisuchus is an extinct monospecific genus of giant, flat-headed crocodyliform within the family Aegyptosuchidae. It was found in the Kem Kem Formation of southeast Morocco, which dates back to the Cenomanian age of the Late Cretaceous epoch. The type species Aegisuchus witmeri was named in 2012 by paleontologists Casey Holliday and Nicholas Gardner, who nicknamed it "Shieldcroc" for the shield-like shape of its skull. A. witmeri is known from a single partial skull including the braincase and skull roof.
Dyrosaurus
Dyrosaurus is a genus of extinct crocodylomorph that lived during the Early Eocene. The name Dyrosaurus comes from () the Greek for lizard or reptile, and Dyr for Djebel Dyr (mountain) close to where the type species was discovered. It was a large reptile with an estimated body length of .
Guarinisuchus
thumb|left|Size (2), compared to other Brazilian Cretaceous Crocodylomorphs
Razanandrongobe
Razanandrongobe (meaning "ancestor [of the] large lizard" in Malagasy) is a genus of carnivorous ziphosuchian crocodyliform from the Middle Jurassic of Madagascar. It contains the type and only species Razanandrongobe sakalavae, named in 2004 by Simone Maganuco and colleagues based on isolated bones found in 2003. The remains, which included a fragment of maxilla and teeth, originated from the Bathonian-aged Sakaraha Formation of Mahajanga, Madagascar. While they clearly belonged to a member of the Archosauria, Maganuco and colleagues refrained from assigning the genus to a specific group beca