Prehistoric therapsid genera

Inostrancevia is an extinct genus of large carnivorous therapsids which lived during the Late Permian in what is now European Russia and Southern Africa. The first-known fossils of this gorgonopsian were discovered in the context of a long series of excavations carried out from 1899 to 1914 in the Northern Dvina, Russia. Among these are two near-complete skeletons embodying the first described specimens of this genus, being also the first gorgonopsian identified in Russia. Several other fossil materials were discovered there, and the various finds led to confusion as to the exact number of val

Gorgonops (from 'Gorgon' and 'eye, face', literally 'Gorgon eye' or 'Gorgon face') is an extinct genus of gorgonopsian therapsid, of which it is the type genus. Gorgonops lived during the Late Permian (Wuchiapingian), about 260–254 million years ago in what is now South Africa.

Moschops (Greek for "calf face") is an extinct genus of therapsids that lived in the Guadalupian epoch, around 265–260 million years ago. They were heavily built plant eaters, and they may have lived partly in water, as hippopotamuses do. They had short, thick heads and might have competed by head-butting each other. Their elbow joints allowed them to walk with a more mammal-like gait rather than crawling. Their remains were found in the Karoo region of South Africa, belonging to the Tapinocephalus Assemblage Zone. Therapsids, such as Moschops, are synapsids, the dominant land animals in the P

Anteosaurus is an extinct genus of large carnivorous dinocephalian synapsid. It lived at the end of the Guadalupian (Middle Permian) during the Capitanian age, about 265 to 260 million years ago, in what is now South Africa. It is mainly known from cranial remains and few postcranial bones. Measuring long and weighing about , Anteosaurus was the largest known carnivorous non-mammalian synapsid and the largest terrestrial predator of the Permian period. Occupying the top of the food chain in the Middle Permian, its skull, jaws and teeth show adaptations to capture large prey, such as giant tita

Titanophoneus ("titanic murderer") is an extinct genus of carnivorous dinocephalian therapsid from the Middle Permian. It is classified within the family Anteosauridae. The type species is Titanophoneus potens. Remains of Titanophoneus have been found at Isheevo, Russia.

Estemmenosuchus (meaning "crowned crocodile" in Greek) is an extinct genus of large, early omnivorous therapsid. It is believed and interpreted to have lived during the middle part of the Middle Permian around 267 million years ago. The two species, E. uralensis and E. mirabilis, are characterised by distinctive horn-like structures, which were probably used for intra-specific display. Both species of Estemmenosuchus are from the Perm (or Cis-Urals) region of Russia. Two other estemmenosuchids, Anoplosuchus and Zopherosuchus, are now considered females of the species E. uralensis. There were m

Lycaenops ("wolf-face") is a genus of carnivorous therapsids. It lived during the Middle Permian to the early Late Permian, about 260 mya, in what is now South Africa.

Titanosuchus ("fierce titan crocodile") is an extinct genus of dinocephalian therapsids that lived in the Middle Permian epoch in South Africa. This genus has only one species, Titanosuchus ferox. Along with its close relatives, Jonkeria and Moschops, Titanosuchus inhabited present-day South Africa around 265 million years ago, in the Late Permian. Titanosuchus is frequently cited as being a carnivore; however, this is based on specimens now assigned to Anteosaurus. Instead, Titanosuchus was likely an omnivorous or herbivorous animal like the related Jonkeria. Titanosuchus is known from fragme

Tapinocephalus ("low, depressed head") is an extinct genus of large herbivorous dinocephalians that lived during the Middle Permian Period in what is now South Africa. Only the type species, Tapinocephalus atherstonei is now considered valid for this genus.

Sauroctonus ("lizard killer") is an extinct genus of gorgonopsian therapsids who lived during the end of the Middle Permian in what is now European Russia. The first fossils, discovered in Tatarstan, were initially believed to belong to a new species of the South African genus Arctognathus (named A. progressus in 1938). The taxon was designated as such until 1940, when it was assigned to the genus Inostrancevia by Ivan Yefremov, before being definitively classified in a separate genus erected by Alexey Bystrow (in 1955). The most complete, known fossils of S. progressus include cranial and pos

Eotitanosuchus ("dawn giant crocodile") is an extinct genus of biarmosuchian therapsids whose fossils were found in the town of Ochyor in Perm Krai, Russia. It lived about 267 million years ago. The only species is Eotitanosuchus olsoni.

Ulemosaurus is an extinct genus of dinocephalian therapsids that lived 265 to 260 million years ago, at Isheevo in Russian Tatarstan. It was a tapinocephalid, a group of bulky herbivores which flourished in the Middle Permian. Ulemosaurus and other tapinocephalians disappeared at the end of the Middle Permian.

Deuterosaurus is an extinct genus of dinocephalian therapsids, one of the non-mammalian synapsids dominating the land during the late Paleozoic.

Dinogorgon is a genus of gorgonopsid from the Late Permian of South Africa and Tanzania. The generic name Dinogorgon is derived from Greek, meaning "terrible gorgon", while its species name rubidgei is taken from the surname of renowned Karoo paleontologist, Professor Bruce Rubidge, who has contributed to much of the research conducted on therapsids of the Karoo Basin. The type species of the genus is D. rubidgei.

Arctops ("bear face") is an extinct genus of gorgonopsian therapsids known from the Late Permian of South and Eastern Africa. It measured up to in length and its skull was long. The type species is Arctops willistoni, named from a poorly prepared and incomplete skull. A second species was named A. watsoni based on a complete skull in 1953, followed by a third in 1970, A. kitchingi, from a smaller, juvenile skull. Both were formally synonymized with A. willistoni by Christian Kammerer in 2017. An additional species, A. umulunshi, was described in 2025 from the Madumabisa Mudstone Formation of Z

Jonkeria is an extinct genus of dinocephalians. Jonkeria was a large and omnivorous animal, from the Tapinocephalus Assemblage Zone, Lower Beaufort Group, of the South African Karoo.

Biarmosuchus is an extinct genus of biarmosuchian therapsids that lived around 267 mya during the Middle Permian period. Biarmosuchus was discovered in the Perm region of Russia. The first specimen was found in channel sandstone that was deposited by flood waters originating from the young Ural Mountains.

Rubidgea is a genus of gorgonopsian from the upper Permian of South Africa and Tanzania, containing the species Rubidgea atrox. The generic name Rubidgea is sometimes believed to be derived from the surname of renowned Karoo paleontologist, Professor Bruce Rubidge, who has contributed to much of the research conducted on therapsids of the Karoo Basin. However, this generic name was actually erected in honor of Rubidge's paternal grandfather, Sidney Rubidge, who was a renowned fossil hunter. Its species name atrox is derived from Latin, meaning "fierce, savage, terrible". Rubidgea is part of th

Syodon (from , "close, shut" and , "tooth", "closed-root tooth") is an extinct genus of dinocephalian therapsids that lived approximately 267-260 million years ago during the middle Permian period of the Paleozoic era. These therapsids, discovered in Russia were initially believed to be true mammals. Syodon was first named by Stephan Kutorga in 1838.

thumb|left|Restoration of H. boonstrai Hipposaurus ('horse lizard') is an extinct genus of basal therapsids known from the Tapinocephalus Assemblage Zone of the Main Karoo Basin, South Africa. Chronologically this is within the Capitanian stage of the Guadalupian Series (Middle Permian). The genus was first described by S.H. Haughton as H. boonstrai on the basis of a skull and associated skeleton and was later considered a gorgonopsian in the family 'Ictidorhinidae' by Robert Broom. It is now considered a basal biarmosuchian, but its affinities remain uncertain. Some authors note the similari

thumb|left|Life restoration of C. longiceps Cyonosaurus is a genus of gorgonopsian therapsids from the late Permian of South Africa. Some skulls have been reported from Early Triassic strata, but further investigation revealed that these reports were erroneous. Cyonosaurus was in length, with a skull in length. The type species Cyonosaurus longiceps was named in 1937.

Aelurognathus is an extinct genus of gorgonopsian therapsids from the Permian of South Africa and Zambia.

Aloposaurus is an extinct genus of gorgonopsian therapsids from the Late Permian of South Africa. It was first named by Robert Broom in 1910, and contains the type species A. gracilis (holotype AMNH 5317), and possibly a second species A. tenuis. This small gorgonopsid had a slender narrow skull only long, with a total body length of .

Phthinosuchus is an extinct genus of therapsids from the Middle Permian of Russia. Phthinosuchus is the sole member of the family Phthinosuchidae. Phthinosuchus may have been one of the most primitive therapsids, meaning that its ancestors may have branched off early from the main therapsid line.

thumb|left|Restoration of S. laticeps with prey thumb|left|Holotype of S. nowaki

thumb|left|Styracocephalus|253x253px Styracocephalus platyrhynchus (Greek for "spiked-head") is an extinct genus of dinocephalian therapsid that existed during the mid-Permian throughout South Africa, but mainly in the Karoo Basin. It is often referred to by its single known species Styracocephalus platyrhynchus. The Dinocephalia clade consisted of the largest land vertebrates and herbivores during the early to mid-Permian. This period is often also referred to as the Guadalupian epoch, approximately 270 to 260 million years ago.

Viatkogorgon is a genus of gorgonopsian (a type of therapsid, the group that includes modern mammals) that lived during the Permian period in what is now Russia. The first fossil was found at the Kotelnich locality near the Vyatka River and was made the holotype of the new genus and species V. ivakhnenkoi in 1999. The generic name refers to the river and the related genus Gorgonops—the gorgons of Greek mythology are often referenced in the names of the group. The specific name honors the paleontologist Mikhail F. Ivakhnenko. The holotype skeleton is one of the most complete gorgonopsian specim

Arctognathus is an extinct genus of gorgonopsids that throve during the Late Permian in the Karoo basin of what is now South Africa.

Archaeosyodon is an extinct genus of dinocephalian therapsids. It was medium-sized, reaching about 1.5–2 m (4–5 ft) in length.

Herpetoskylax is an extinct genus of biarmosuchian therapsids which existed in South Africa. The type species is Herpetoskylax hopsoni. It lived in the Late Permian Period.

Keratocephalus ("horned head") is an extinct genus of tapinocephalian therapsids from the early Capitanian age of South Africa.

Niaftasuchus is an extinct genus of therapsids. Its type and only named species is Niaftasuchus zekkeli.

Aelurosaurus ("cat lizard", from Ancient Greek "cat" and "lizard") is a small, carnivorous, extinct genus of gorgonopsian therapsids from the Late Permian of South Africa. It was discovered in the Karoo Basin of South Africa, and first named by Richard Owen in 1881. It was named so because it appeared to be an ancestor for cat-like marsupials, but not yet a mammal itself. It contains five species, A. felinus, A. whaitsi, A. polyodon, A. wilmanae, and A.? watermeyeri. A. felinus, the type species, is generally well described with established features, while the other four species are not due to

Proburnetia is an extinct genus of biarmosuchian therapsids in the family Burnetiidae, from the Late Permian of Russia. It had bizarre bumps and protrusions on its skull.

Scylacops (meaning "face that tears") is an extinct genus of Gorgonopsia. It was first named by Broom in 1913, and contains two species, S. bigendens, and S. capensis. Its fossils have been found in South Africa and Zambia. It is believed to be closely related to the Gorgonopsian Sauroctonus progressus. Scylacops was a moderately sized Gorgonopsid.

Ictidorhinus is an extinct genus of biarmosuchian therapsids. Fossils have been found from the Dicynodon Assemblage Zone of the Beaufort Group in the Karoo Basin, South Africa and are of Late Permian age. It had a short snout and proportionally large orbits. These characteristics may be representative of a juvenile animal, possibly of Lycaenodon. However, these two genera are not known to have existed at the same time, making it unlikely for Ictidorhinus material to be from a juvenile form of Lycaenodon.

Paraburnetia is an extinct genus of biarmosuchian therapsids from the Late Permian of South Africa. It is known for its species P. sneeubergensis and belongs to the family Burnetiidae. Paraburnetia lived just before the Permian–Triassic mass extinction event.

Australosyodon is an extinct genus of dinocephalian therapsids from the middle Permian of South Africa. The first fossil was discovered in the 1980s near the village of Prince Albert Road in the Karoo region of South Africa.

Pravoslavlevia is an extinct genus of gorgonopsian therapsids that lived in the late Permian and is part of the Sokolki subcomplex of Russia. It had a skull long. The total length of the animal was about . Only one species (P. parva) is known. ==Classification== thumb|Pravoslavlevia The following cladogram showing the position of Pravoslavlevia within Gorgonopsia follows Kammerer and Masyutin, 2018:

Pampaphoneus is an extinct genus of carnivorous dinocephalian therapsid belonging to the family Anteosauridae. It lived 268 to 265 million years ago during the Wordian age of the Guadalupian (= middle Permian) period in what is now Brazil. Pampaphoneus is known by an almost complete skull with the lower jaw still articulated, discovered on the lands of the Boqueirão Farm, near the city of São Gabriel, in the state of Rio Grande do Sul. A second specimen from the same locality was reported in 2019 and 2020 but has not yet been described. It is composed of a skull associated with postcranial rem

Bullacephalus is an extinct genus of biarmosuchian therapsids belonging to the family Burnetiidae. The type species B. jacksoni was named in 2003. It is known from a relatively complete skull and lower jaw, discovered in the Late Permian Tapinocephalus Assemblage Zone of the Beaufort Group of South Africa. This genus of therapsida lived during the Late Permian period, approximately 250 million years ago.

Rhopalodon is an extinct genus of therapsids from the Permian of Russia. It has been variously classified as a dinosaur, a dinocephalian, or another branch of amniotes. Rhopalodon is notable for being among the first "reptiles" mentioned in Nature. T.H. Huxley wrote of this animal, among others, in the inaugural issue of the magazine, in November 1869. He gave the age of this animal and of the contemporary Deuterosaurus as Triassic, but both are now known to have lived during the Middle Permian.

Burnetia is an extinct genus of biarmosuchian therapsids in the family Burnetiidae, from the Late Permian of South Africa. Burnetia is known so far from a single holotype skull lacking the lower jaws described by South African paleontologist Robert Broom in 1923. Due to erosion and dorsoventral crushing, features of the skull are hard to interpret. Stutural lines are further distorted by the unusual shape of the skull roof, including many bosses and protuberances.

thumb|left|Restoration of C. rubidgei thumb|left|Referred skull, the holotype of Tigrisaurus pricei

Brithopus is an extinct genus of dinocephalian therapsids. It contains a single species, Brithopus priscus, known from fragmentary remains found in the Copper Sandstones near Isheevo, Russia.

Nochnitsa is an extinct genus of gorgonopsian therapsids who lived during an uncertain stage of the Permian in what is now European Russia. Only one species is known, N. geminidens, described in 2018 from a single specimen including a complete skull and some postcranial remains, discovered in the red beds of Kotelnich, Kirov Oblast. The genus is named in reference to Nocnitsa, a nocturnal creature from Slavic mythology. This name is intended as a parallel to the Gorgons, which are named after many genera among gorgonopsians, as well as for the nocturnal behavior inferred for the animal. The on

Kamagorgon is an extinct genus of therapsids from the Middle Permian of Russia. The type and only species is Kamagorgon ulanovi. It is only known from an incomplete skull. The snout is short and the canine teeth are very large. Kamagorgon was named in 1998 and originally classified in the biarmosuchian family Eotitanosuchidae along with the poorly known therapsid Eotitanosuchus. More recently, Kamagorgon has considered as a primitive gorgonopsian rather than a biarmosuchian due to the length of the front jawbone and rear side of the skull. These features are commonly shared by the brithopodid

Sinophoneus is an extinct genus of carnivorous dinocephalian therapsid belonging to the family Anteosauridae. It lived 272 to 270 million years ago at the beginning of the Middle Permian (Lower Roadian) in what is now the Gansu Province in northern China. It is known by a skull of an adult individual (the holotype GMV1601), as well as by many skulls of juvenile specimens. The latter were first considered as belonging to a different animal, named Stenocybus, before being reinterpreted as immature Sinophoneus. Sinophoneus shows a combination of characters present in other anteosaurs. Its bulbous

Raranimus is an extinct genus of therapsids of the Middle Permian. It was described in 2009 from a partial skull found in 1998 from the Dashankou locality of the Qingtoushan Formation, outcropping in the Qilian Mountains of Gansu, China. The genus is the most basal known member of the clade Therapsida, to which the later Mammalia belong.

Ivantosaurus is an extinct genus of therapsid that lived in Russia during the Wordian stage of the Permian period, named in honor of paleontologist Ivan Antonovich Efremov.

Lemurosaurus is a genus of extinct biarmosuchian therapsids from the Late Permian of South Africa. The generic epithet Lemursaurus is a mix of Latin, lemures "ghosts, spirits", and Greek, sauros, "lizard". Lemurosaurus is easily identifiable by its prominent eye crests, and large eyes. The name Lemurosaurus pricei was coined by paleontologist Robert Broom in 1949, based on a single small crushed skull, measured at approximately 86 millimeters in length, found on the Dorsfontein farm in Graaff-Reinet. To date, only two skulls of the Lemurosaurus have been discovered, so body size is unknown. Th

Notosyodon is an extinct genus of non-mammalian therapsids. The holotype PIN 2505/1, consists of a partial skull preserving the orbital, occipital, and basicranial regions. Other remains include PIN 2505/2, a right lower incisor, and PIN 2505/3, a left upper postcanine, found associated with the holotype and PIN 2608/1, the anterior half of a left dentary found on the right bank of the Donguz River, near Dolmatovskii Farm, Sol-Iletsk District, Orenburg Region, Russia.

Nikkasaurus is an extinct genus of therapsids first named and described by Ivakhnenko.

Pachydectes is an extinct genus of biarmosuchian therapsids from the Middle Permian of South Africa known from a single skull. The etymology of the name Pachydectes is derived from the Greek word pakhus, meaning "thick" or "thickened", and dektes, meaning "biter". In conjunction this name is representative of the unique pachyostotic bone present above the maxillary canine tooth found in the skull of the specimen. There is only one known species within the genus, Pachydectes elsi which is named in honor of the person who discovered the fossil.

Tapinocaninus (Greek for "humble"- tapino, and "canine"- caninus) is an extinct genus of therapsids in the family Tapinocephalidae, of which it is the most basal member. Only one species is known, Tapinocaninus pamelae (meaning "Pamela's humble canine"). The species is named in honor of Rubidge's mother, Pamela. Fossils have been found dating from the Middle Permian (Wordian age).

Struthiocephalus ("ostrich head") is an extinct genus of dinocephalian therapsids from the Permian of South Africa. It was a large animal, reaching in body mass.

Smilesaurus is an extinct genus of gorgonopsian known from South Africa. It lived during the Late Permian. It contains the single species S. ferox.

Microurania is an extinct genus of therapsids from the Middle Permian first named and described by Mikhaïl Ivakhnenko. It is known from a single partial skull found in the region of Orenburg, Russia. According to Kammerer, 2011, it likely represents the remains of a juvenile dinocephalian.

Eriphostoma is an extinct genus of gorgonopsian therapsids known from the Middle Permian (middle Capitanian stage) of Tapinocephalus Assemblage Zone, South Africa. It has one known species, Eriphostoma microdon, and was first named by Robert Broom in 1911. It is one the oldest known gorgonopsian and among the smallest and most basal members of the clade.

Phthinosaurus is an extinct genus of therapsids from the Middle Permian of Russia. The type species Phthinosaurus borrisiaki was named by Soviet paleontologist Ivan Yefremov in 1940 on the basis of an isolated lower jaw. Because this jaw provides few distinguishing characteristics, the evolutionary relationships of Phthinosaurus are poorly known. Yefremov named the family Phthinosuchidae in 1954 to include Phthinosaurus and the newly named Phthinosuchus, which was described on the basis of a crushed partial skull. American paleontologist Everett C. Olson placed both of these therapsids in the