Cambrian first appearances

The annelids (), also known as the segmented worms, are animals that comprise the phylum Annelida (; ). The phylum contains over 22,000 extant species, including ragworms, earthworms, and leeches. The species exist in and have adapted to various ecologies – some in marine environments as distinct as tidal zones and hydrothermal vents, others in fresh water, and yet others in moist terrestrial environments.

Trilobites (; meaning "three-lobed entities") are extinct marine arthropods that form the class Trilobita. One of the earliest groups of arthropods to appear in the fossil record, trilobites were among the most successful of all early animals, existing in oceans for almost 270million years, with over 22,000 species having been described. Because trilobites had wide diversity and an easily fossilized mineralised exoskeleton made of calcite, they left an extensive fossil record. The study of their fossils has facilitated important contributions to biostratigraphy, paleontology, evolutionary biol

Portion of the Cambrian Period during which life vastly diversified

Graptolites are a group of colonial animals, members of the subclass Graptolithina within the class Pterobranchia. These filter-feeding organisms are known chiefly from fossils found from the Middle Cambrian (Miaolingian, Wuliuan) through the Lower Carboniferous (Mississippian). A possible early graptolite, Chaunograptus, is known from the Middle Cambrian. Recent analyses have favored the idea that the living pterobranch Rhabdopleura represents an extant graptolite which diverged from the rest of the group in the Cambrian.

Leptostraca (from the Greek words for thin and shell) is an order of small, marine crustaceans. Its members, including the well-studied Nebalia, occur throughout the world's oceans and are usually considered to be filter-feeders. It is the only extant order in the subclass Phyllocarida. They are believed to represent the most primitive members of their class, the Malacostraca, and first appear in the fossil record during the Cambrian period.

The Rostroconchia is a class of extinct molluscs dating from the early Cambrian to the Late Permian. They were initially thought to be bivalves, but were later given their own class. They have a single shell in their larval stage, and the adult typically has a single, pseudo-bivalved shell enclosing the mantle and muscular foot. The anterior part of the shell probably pointed downward and had a gap from which the foot could probably emerge. Rostroconchs probably lived a sedentary semi-infaunal lifestyle. There were probably more than 1,000 species of members of this class.

Hyoliths are an extinct group of invertebrates with small conical shells, known from fossils from the Palaeozoic era. They are at least considered lophotrochozoans, possibly being lophophorates, a group which includes the brachiopods (hyoliths may even be brachiopods themselves), while others consider them as being basal lophotrochozoans, or even molluscs.

Dinocaridida is a proposed fossil taxon of basal arthropods, which flourished during the Cambrian period and survived up to Early Devonian. Characterized by a pair of frontal appendages and series of body flaps, the name of Dinocaridids (Greek for deinos "terrible" and Latin for caris "crab") refers to the suggested role of some of these members as the largest marine predators of their time. Dinocaridids are occasionally referred to as the 'AOPK group' by some literatures, as the group composed of Radiodonta (Anomalocaris and relatives), Opabiniidae (Opabinia and relatives), and the "gilled lo

Agnostida are an order of extinct arthropods which have classically been seen as a group of highly modified trilobites, though some recent research has doubted this placement. Regardless, they appear to be close relatives as part of the Artiopoda. They are present in the Lower Cambrian fossil record along with trilobites from the Redlichiida, Corynexochida, and Ptychopariida orders, and were highly diverse throughout the Cambrian. Agnostidan diversity severely declined during the Cambrian-Ordovician transition, and the last agnostidans went extinct in the Late Ordovician.

Naraoia is a genus of small-to-average-size (about long) marine arthropods within the family Naraoiidae, that lived from the early Cambrian to the late Silurian period. The species are characterized by a large alimentary system and sideways oriented antennas.

Edrioasteroidea is an extinct class of echinoderms. The living animal would have resembled a pentamerously symmetrical disc or cushion. They were obligate encrusters and attached themselves to inorganic or biologic hard substrates (frequently hardgrounds or brachiopods). A 507 million years old species, Totiglobus spencensis, is actually the first known echinoderm adapted to live on a hard surface after the soft microbial mats that covered the seafloor were destroyed in the Cambrian substrate revolution.

Asaphida is a large, morphologically diverse order of trilobites found in marine strata dated from the Middle Cambrian until their extinction during the Silurian. Asaphida contains six superfamilies (Anomocaroidea, Asaphoidea, Cyclopygoidea, Dikelocephaloidea, Remopleuridoidea and Trinucleioidea), but no suborders. Asaphids comprise some 20% of described fossil trilobites.

The Eocrinoidea were an extinct class of echinoderms that lived between the Early Cambrian and Late Silurian periods. They are the earliest known group of stalked, brachiole-bearing echinoderms, and were the most common echinoderms during the Cambrian.

Ptychopariida is a large, heterogeneous order of trilobite containing some of the most primitive species known. The earliest species occurred in the second half of the Lower Cambrian, and the last species did not survive the Ordovician–Silurian extinction event.

The halkieriids are a group of fossil organisms from the Lower to Middle Cambrian. Their eponymous genus is Halkieria .

Thylacocephala (from the Greek or ', meaning "pouch", and or ' meaning "head") is an extinct group of mandibulate arthropods, that are generally regarded as a kind of crustacean, though their exact position within this group is uncertain. As a class they have a short research history, having been erected in the early 1980s.

Orthida is an extinct order of brachiopods which appeared during the Early Cambrian period and became very diverse by the Ordovician, living in shallow-shelf seas. Orthids are the oldest member of the subphylum Rhynchonelliformea (Articulate Brachiopods), and is the order from which all other brachiopods of this group stem.

Harpetida is one of the eleven orders of the extinct arthropod class Trilobita. The first harpetid trilobites appear in the Upper Cambrian, and the last species die out in the late Devonian period.

Myllokunmingiidae is a family of primitive jawless fishes which lived during the Cambrian period. The myllokunmingiids are considered to be the earliest known vertebrate animals. The group contains only three known genera, Haikouichthys, Myllokunmingia, and Zhongjianichthys. Their fossils have been found only in the Maotianshan Shales lagerstätte.

Nektaspida (also called Naraoiida, Nektaspia and Nectaspida) is an extinct order of non-mineralised artiopodan arthropods. They are known from the lower-Cambrian to the upper Silurian. Originally classified as trilobites, which they superficially resemble, they are now placed as close relatives as members of the Trilobitomorpha within Artiopoda. The order is divided into three major families; Emucarididae, Liwiidae, and Naraoiidae.

Lingulella is a genus of phosphatic-shelled brachiopod. It is known from the Middle Cambrian Burgess Shale (Canada) to the Upper Ordovician Bromide Formation (United States) in North America. 346 specimens of Lingulella are known from the Greater Phyllopod bed, where they comprise 0.66% of the community.

Vauxia is an extinct genus of demosponge that had a distinctive branching mode of growth. Each branch consisted of a network of strands. Vauxia also had a skeleton of spongin (flexible organic material) common to modern day sponges. Much like Choia and other sponges, Vauxia fed by extracting nutrients from the water.

Pentamerida is an order of biconvex, impunctate shelled, articulate brachiopods that are found in marine sedimentary rocks that range from the Middle Cambrian through the Devonian.

Choia is a genus of extinct demosponge ranging from the Cambrian until the Lower Ordovician periods. Fossils of Choia have been found in the Burgess Shale in British Columbia; the Maotianshan shales of China; the Wheeler Shale in Utah; the Itajaí Basin in Brazil; and the Lower Ordovician Fezouata formation. It was first described in 1920 by Charles Doolittle Walcott.

Redlichiina is a suborder of the order Redlichiida of trilobites. The suborder contains three superfamilies: Emuelloidea, Redlichioidea and Paradoxidoidea. These trilobites are some of the oldest trilobites known. They originated at the beginning of the Cambrian Period and disappeared (possibly by evolving into members of the Ptychopariida order) at the end of the middle Cambrian.

Agnostidae is a family of Agnostida trilobites. Like all Agnostina, they were eyeless and had only two thoracic segments. These trilobites inhabited benthic waters worldwide from 508 to 461 million years ago. The family includes the following genera, among others:

Hymenocarina is an order of extinct marine arthropods known from the Cambrian. They possess bivalved carapaces, typically with exposed posteriors. Members of the group are morphologically diverse and had a variety of ecologies, including as filter feeders and as predators. Recent research has generally considered them to be stem or crown group members of Mandibulata, due to the presence of mandibles in well-preserved species.

Yukoniidae S. Zhang, 1980 [nom. transl. et emend. Jell, in Whittington et al., 1997 ex Yukoniinae S. Zhang in W. Zhang, Lu et al., 1980] is a small family of trilobites, belonging to the Eodiscina.

Artiopoda is a clade of extinct arthropods that includes trilobites and their close relatives. It was erected by Hou and Bergström in 1997 to encompass a wide diversity of arthropods that would traditionally have been assigned to the Trilobitomorpha. Trilobites, in part due to abundance of findings owing to their mineralized exoskeletons, are by far the best recorded, diverse, and long lived members of the clade. Other members, which lack mineralised exoskeletons, are known mostly from Cambrian deposits.

The ”Fallotaspidoidea” are a superfamily of trilobites, a group of extinct marine arthropods. It lived during the Lower Cambrian (Atdabanian) and species occurred on all paleocontinents except for the Gondwana heartland (currently Latin America, most of Africa, Australia, Antarctica, India and China). A member of this group, Profallotaspis jakutensis, has long been the earliest known trilobite, but recently the redlichiid Lemdadella has been claimed as occurring even earlier.

The Paradoxididae are a family of trilobites, a group of extinct marine arthropods. They occurred during the late Lower Cambrian (Toyonian) and disappeared at the end of the Middle Cambrian. Representatives of this family have been found in the paleocontinents of Avalonia, Baltica, and Gondwana, now Canada (Nova-Scotia, New Brunswick, Newfoundland), USA (Massachusetts, South Carolina), England, Wales, Morocco, Spain, Czech Republic, Poland, Russia (Novaya Zemlya, Northern Siberia, North-East Yakutia), Mongolia, and Turkey. Species in this family can typically grow large to very large (over 30&

Hazelia is a genus of spicular Cambrian demosponge known from the Burgess Shale, the Marjum formation of Utah, and possibly Chengjiang. It was described by Charles Walcott in 1920.

Eodiscidae is a family of agnostid trilobites that lived during the final Lower Cambrian (late Toyonian) and the Middle Cambrian. They are small or very small, and have a thorax of two or three segments. Eodiscidae includes nine genera (see box).

Asaphidae is a family of asaphid trilobites. Although the first genera originate in Upper Cambrian marine strata, the family becomes the most widely distributed and most species-rich trilobite family during the Ordovician.

The palaeoscolecids are a group of extinct ecdysozoan worms resembling armoured priapulids. They are known from the Lower Cambrian to the lower Ludfordian (late Silurian); they are mainly found as disarticulated sclerites, but are also preserved in many of the Cambrian lagerstätten. They take their name from the typifying genus Palaeoscolex. Other genera include Cricocosmia from the Lower Cambrian Chengjiang biota. Their taxonomic affinities within Ecdysozoa have been the subject of debate.

Redlichiidae is a family of redlichiid trilobites which lived from the Botomian to the Middle Cambrian period. It contains the following genera, divided between five subfamilies:

Liwiidae is a family of arthropods in the order Nektaspida. Members are known from the Cambrian and Ordovician periods.

Vicissicaudata is an unranked group of artiopodan arthropods, containing Cheloniellida, the speciose Aglaspidida, and several other genera outside these groups like Sidneyia and Emeraldella.

The Paradoxidoidea Hawle & Corda 1847, are a superfamily of trilobites, a group of extinct marine arthropods. They occurred during the late Lower Cambrian (Toyonian) and disappeared at the end of the Middle Cambrian.

Olenellidae is an extinct family of redlichiid trilobite arthropods. Olenellids lived during the late Lower Cambrian (Botomian/Toyonian) in the Olenellus-zone in the former paleocontinent of Laurentia and parts of what became the Famatinian orogen in what is now Argentina. This family can be distinguished from most other Olenellina by the partial merger of the frontal (L3) and middle pair (L2) of lateral lobes of the central area of the cephalon, that is called glabella, creating two isolated slits.

Markuelia is a genus of fossil worm-like bilaterian animals allied to Ecdysozoa and known from strata of Lower Cambrian to Lower Ordovician age containing five species.