Enicocephalomorpha

Enicocephalidae, also called unique-headed bugs and gnat bugs, are a family of around 300 species of the suborder Heteroptera. They are typically long, and found throughout the world. They have an elongated head, constricted in places, hence their head is 'unique'. left|thumb|Systelloderes maclachlani They are classified into about 47 genera placed in five subfamilies. The family members can be separated from those of the Aenictopecheidae on the basis of the pronotal division into three lobes (except in the genus Alienates). They also show polymorphism with winged males and wingless or short-w

Aenictopecheidae is a rare family of insects occurring worldwide but containing only a few species, including a single American species, Boreostolus americanus. This species lives under large, flat stones and sandy substrates along mountain streams in Oregon, Washington, and Colorado. It is 5 mm long and occurs in both the macropterous and brachypterous condition. It is assumed to be predaceous.

Enicocephaloidea is a sole superfamily within the infraorder Enicocephalomorpha of the hemipteran suborder Heteroptera ("true bugs"). The group was formerly thought to be related to the Reduviidae because of similarities in head structure but they are now considered different enough from other Heteroptera to constitute a separate infraorder and represents a sister group of the Leptopodomorpha. They are predaceous and some species are known to swarm but little is known of their life history. A few species of enicocephalids are known to shed their wings before entering the surface of subsoil.

Systelloderes is a genus of gnat bugs in the family Enicocephalidae. There are about 12 described species in Systelloderes.