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Huayquerian

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Caenolestidae
family of mammals
Macrauchenia
Macrauchenia ("long llama", based on the now-invalid llama genus, Auchenia, from Greek "big neck") is an extinct genus of large ungulate native to South America from the Pliocene or Middle Pleistocene to the end of the Late Pleistocene. It was one of the last surviving members of the extinct order Litopterna, a group of South American native ungulates distinct from the two orders which contain all living ungulates which had been present in South America since the early Cenozoic, over 60 million years ago, prior to the arrival of living ungulates in South America around 2.5 million years ago as
Thylacosmilus
Thylacosmilus is an extinct genus of saber-toothed metatherian mammals that inhabited South America from the Late Miocene to Pliocene epochs. Though Thylacosmilus looks similar to the "saber-toothed cats", it was not a felid, like the well-known American Smilodon, but a sparassodont, a group closely related to marsupials, and only superficially resembled other saber-toothed mammals due to convergent evolution, with the aforementioned Thylacosmilus being one of the last known sparassodonts. A 2005 study found that the bite forces of Thylacosmilus and Smilodon were low, which indicates that the
Thylamys
Thylamys is a genus of opossums in the family Didelphidae. The premaxillae are rounded rather than pointed. The females lack a pouch. The females' nipples are arranged in two symmetrical rows on the abdomen. All species but T. macrurus store fat in their tails, although this is not necessarily true for all species in the genus. Fossils belonging to the genus date back to the Miocene, with the oldest specimens being found in the Cerro Azul Formation of Argentina and the Honda Group of Colombia. Genetic studies indicate that the genus may have originated around 14 million years ago.
Toxodon
Toxodon (from Ancient Greek τόξον (tóxon), meaning "bow", and ὀδούς (odoús), meaning "tooth", in reference to the curvature of the teeth) is an extinct genus of large ungulate native to South America from the Pliocene to the end of the Late Pleistocene. Toxodon is a member of Notoungulata, an order of extinct South American native ungulates distinct from the two living ungulate orders that had been indigenous to the continent for over 60 million years since the early Cenozoic, prior to the arrival of living ungulates into South America around 2.5 million years ago during the Great American Int
Purussaurus
Purussaurus is a genus of extinct giant caimans that lived in the Americas during the Miocene epoch, from the Friasian to the Huayquerian in the SALMA classification. It is known from skull material found in the Brazilian and Peruvian Amazon, Argentina, Colombian Villavieja Formation, Panamanian Culebra Formation, Urumaco, and Socorro Formations of northern Venezuela.
Stupendemys
Stupendemys is an extinct genus of freshwater side-necked turtle, belonging to the family Podocnemididae. It is the largest freshwater turtle known to have existed, with a carapace over 2 meters long. Its fossils have been found in northern South America, in rocks dating from the Middle Miocene to the very start of the Pliocene, about 13 to 5 million years ago. Male specimens are known to have possessed bony horns growing from the front edges of the shell and the discovery of the fossil of a young adult shows that the carapace of these turtles flattens with age. A fossil skull described in 202
Thalassocnus
Thalassocnus is an extinct genus of semiaquatic ground sloths from the Miocene and Pliocene of the Pacific South American coast. It is monotypic within the subfamily Thalassocninae. The five species—T. antiquus, T. natans, T. littoralis, T. carolomartini, and T. yuacensis—represent a chronospecies, a population gradually adapting to marine life in one direct lineage. They are the only known aquatic sloths, but they may have also been adapted to a terrestrial lifestyle. They have been found in the Pisco Formation of Peru, the Tafna Formation of Argentina, and the Bahía Inglesa, Coquimbo, and Ho
Andalgalornis
Andalgalornis is a genus of flightless predatory birds of the extinct family Phorusrhacidae (often called "terror birds") that lived in Argentina. The type and only species is A. steulleti.
Pelagornis
Pelagornis is an extinct genus of prehistoric pseudotooth birds, a group of extinct seabirds. Species span from the Oligocene to the Early Pleistocene. Members of Pelagornis represent among the largest pseudotooth birds, with one species, P. sandersi, having the widest wingspan of any bird known.
Argentavis
Argentavis is an extinct genus of teratornithid known from three sites in the Epecuén and Andalhualá Formations in central and northwestern Argentina dating to the Late Miocene (Huayquerian). The type species, A. magnificens, is sometimes called the giant teratorn. Argentavis was among the largest flying birds to ever exist, holding the record for heaviest flying bird, although it was surpassed in wingspan after the 2014 description of Pelagornis sandersi, which is estimated to have possessed wings some 20% longer than those of Argentavis.
Gryposuchus
Gryposuchus (meaning "hooked crocodile") is an extinct genus of gavialid crocodilian. Fossils have been found from Argentina, Colombia, Venezuela, Brazil and the Peruvian Amazon. The genus existed during the Miocene epoch (Colhuehuapian to Huayquerian). One recently described species, G. croizati, grew to an estimated length of . Gryposuchus is the type genus of the subfamily Gryposuchinae, although a 2018 study indicates that Gryposuchinae and Gryposuchus might be paraphyletic and rather an evolutionary grade towards the gharial.
Acrophoca
Acrophoca longirostris, also known as the swan-necked seal, is an extinct genus of Late Miocene pinniped. It was thought to have been the ancestor of the modern leopard seal; however, it is now thought to be a species of monk seal.
Phoberomys
thumb | right | alt=A 10cm skull of Phoberomys pattersoni in ventral view | The skull of Phoberomys pattersoni Phoberomys is an extinct genus of rodents. Fossil specimens from the Late Miocene period have been discovered in the Ituzaingó Formation of Argentina, the Solimões Formation of Brazil, the Urumaco Formation at Urumaco in Venezuela, and the Pliocene of Peru.
Phoberomys pattersoni
species of mammal (fossil)
Carcharodon hubbelli
species of fossil shark
Psilopterus
Psilopterus (Greek for "bare wing") is an extinct genus of phorusrhacid ("terror bird") from the Middle Oligocene to possibly the Late Pleistocene of Argentina and Uruguay. Compared to other phorusrhacids, members of the genus are both relatively gracile and diminutive, and include the smallest known species of terror bird: with the head raised P. bachmanni was in height and weighed about , while the largest members of the genus were only about . The birds resemble the modern cariama (Cariama cristata), except with a heavier build and considerably smaller wings. Fossil finds in Uruguay indicat
Hapalops
thumb|left|Life restoration of Hapalops longiceps and Propalaehoplophorus australis thumb|left|Skeleton
Adinotherium
Adinotherium (meaning "not terrible beast") is an extinct genus of toxodontid, large bodied hoofed ungulates which inhabited South America during the Middle to Late Miocene, from 17.5 to 6.8 Ma and existed for approximately , Santacrucian to Huayquerian in the South American land mammal ages (SALMA). Fossils of Adinotherium have been found in the Santa Cruz and Ituzaingó Formations of Argentina and the Chucal and Río Frías Formations of Chile.
Diadiaphorus
Diadiaphorus is an extinct genus of litoptern mammal from the Miocene of Argentina (Ituzaingó, Pinturas, Chiquimil and Santa Cruz Formations) and Bolivia (Nazareno Formation), South America.
Mourasuchus
Mourasuchus is an extinct genus of giant, aberrant caiman from the Miocene of South America. Its skull has been described as duck-like, being broad, flat, and very elongate, superficially resembling Stomatosuchus from the Late Cretaceous.
Protypotherium
Protypotherium is an extinct genus of notoungulate mammals native to South America during the Oligocene and Miocene epochs. A number of closely related animals date back further, to the Eocene. Fossils of Protypotherium have been found in the Deseadan Fray Bentos Formation of Uruguay, Muyu Huasi and Nazareno Formations of Bolivia, Cura-Mallín and Río Frías Formations of Chile, and Santa Cruz, Salicas, Ituzaingó, Aisol, Cerro Azul, Cerro Bandera, Cerro Boleadoras, Chichinales, Sarmiento and Collón Curá Formations of Argentina.
Telicomys
Telicomys is an extinct genus of rodent from the Solimões Formation, Brazil, South America.
Procariama
Procariama is an extinct monotypic genus of phorusrhacid, which lived from the Late Miocene to the Late Pliocene (11-2 million years ago) of Argentina. Fossils of the animal have been found in six places, in the Cerro Azul and Andalhuala Formations. More specifically in the Andagalá department and in the north of the Belén department of the Catamarca province, with a single location in the La Pampa province. The type and only species, Procariama simplex, is the largest member of the subfamily Psilopterinae.
Cyonasua
Cyonasua (meaning "dog-coati" in Greek) is an extinct genus of procyonid from the Late Miocene to Middle Pleistocene of South America. Fossils of Cyonasua have been found in Argentina (Ituzaingó, Epecuén, Huayquerías, Monte Hermoso, Chapadmalal, Maimará, Ensenada, La Playa, Chiquimil, Andalhuala, and Cerro Azul Formations), Bolivia (Tariquía Formation), Uruguay (Camacho Formation), and Venezuela (San Gregorio Formation). The oldest well-dated fossils of Cyonasua are approximately 7.3 million years old. Most fossils of Cyonasua are late Miocene to early late Pliocene (Huayquerian to Chapadmalal
Trigodon gaudryi
Trigodon is an extinct genus of the family Toxodontidae, a large-bodied notoungulate which inhabited South America during the Late Miocene to Early Pliocene (Mayoan to Montehermosan in the SALMA classification), living from 11.61 to 4.0 Ma which existed for approximately . The type species is T. gaudryi. It bore a superficial resemblance to a rhinoceros, in that it had a horn on its forehead, and was one of a few horned notoungulates, including Adinotherium and Leontinia.
Neotamandua
Neotamandua is an extinct genus of anteaters that lived in the Miocene to Pliocene in South America.
Aulophyseter
Aulophyseter is an extinct genus of sperm whales from the subfamily Physeterinae that existed during the Miocene.
Piscogavialis
Piscogavialis is an extinct genus of gryposuchine gavialid crocodylian. Two species have been described, the type species P. jugaliperforatus and P. laberintoensis. Fossils of Piscogavialis have been found from the Mio-Pliocene Pisco Formation of the Sacaco Basin in southern Peru in 1998, where it coexisted with the much smaller gavialid Sacacosuchus.
Ikanogavialis
Ikanogavialis is an extinct genus of gavialid crocodilian. Fossils have been found in the Urumaco Formation in Urumaco, Venezuela and the Solimões Formation of Brazil. The strata from which remains are found are late Miocene in age, rather than Pliocene as was once thought. A possible member of this genus survived into the Late Holocene on Muyua or Woodlark Island in Papua New Guinea.
Xenastrapotherium
alt=Reconstruction of the astrapothere Xenastrapotherium christi. Artwork created with graphite and colored pencils. Author: Edwin Chávez "Disfrasaurio".|thumb|Paleoart of Xenastrapotherium christi. Artwork by Edwin Chávez "Disfrasaurio". Xenastrapotherium is an extinct genus of astrapothere, a type of hoofed herbivorous mammal, native to South America, which lived in the Middle to Late Miocene period, typically during the Laventan stage. It is a member of the family Astrapotheriidae in the subfamily Uruguaytheriinae, large astrapotheres, equipped with a trunk-like nose and protruding teeth, s
Promegatherium
Promegatherium ("before Megatherium") is a genus of prehistoric xenarthrans that lived in Argentina, during the Late Miocene. This genus is regarded as closely related to the later, and more famous genus, Megatherium, hence the reference in the name. The first specimens of Promegatherium were originally described by the biologist Florentino Ameghino in 1883.
Surameryx
Surameryx is an extinct genus of herbivorous artiodactyls originally described as belonging to the extinct family Palaeomerycidae. A single species, S. acrensis, was described from the Late Miocene (between the Mayoan and Huayquerian SALMA, between 11.6 and 5.3 million years ago) of the Madre de Dios Formation, South America. It was originally interpreted as one of the few northern mammals that entered South America before the Pliocene. However, both its identification as a member of the family Palaeomerycidae and claims about its Miocene age were subsequently challenged.
Acrecebus fraileyi
Acrecebus is a prehistoric cebid monkey from the Late Miocene Solimões Formation of Acre State, Brazil and Bolivia. The only species known is A. fraileyi. This genus is closely related to the genus Cebus.
Thylophorops
Thylophorops is an extinct genus of didelphine opossums from the Pliocene of South America. Compared to their close didelphine cousins like the living Philander and Didelphis (and like the still living Lutreolina) opossums, Thylophorops displays specialization towards carnivory, and one species, T. lorenzinii, is the largest known opossum of all time, which could imply a macropredatory role.
Neoepiblemidae
The Neoepiblemidae are an extinct family of hystricognath rodents from South America. The genera Dabbenea and Perumys are now included in Phoberomys. The delineation between Neoepiblemidae and Dinomyidae has historically been unclear, with some genera (such as Phoberomys and Eusigmomys) having varying taxonomic placement. A 2017 study found Phoberomys to be part of the group, while Eusigmomys was found to be part of the Dinomyidae.
Hoffstetterius
Hoffstetterius is an extinct genus of toxodontid notoungulate mammal, belonging to the subfamily Toxodontinae whose remains were discovered in the Middle to Late Miocene (Mayoan to Montehermosan) Mauri Formation in the La Paz Department in Bolivia. The only described species is the type Hoffstetterius imperator.
Ribodon
Ribodon is an extinct genus of manatee that lived around South America (Ituzaingó Formation, then described as Entrerriana Formation, Argentina, Solimões Formation, Brazil and Urumaco Formation, Urumaco, Venezuela) during the Tortonian (Mayoan to Huayquerian in the South American land mammal ages). The type species is R. limbatus.
Xotodon
Xotodon is an extinct genus of toxodontid mammal that lived during the Late Miocene (Huayquerian in the SALMA classification) in Argentina, South America. Fossils of Xotodon have been found in the Ituzaingó, Maimará and Chiquimil Formations of Argentina.
Eleutherocercus
Eleutherocercus was a genus of glyptodonts that lived during the Late Miocene and Early Pliocene in South America. Fossils of the genus have been found in the Huayquerian Ituzaingó Formation (E. paranensis) and the Montehermosan Monte Hermoso Formation (E. antiquus) in Argentina.
Piscobalaena
Piscobalaena is an extinct genus of cetaceans, which lived from the Middle to Late Miocene epochs (about 11.6 to 5.3 million years ago) in Peru and Florida. Its fossils have been found in the Pisco Formation of Peru and the Bone Valley Formation of Florida. At least some individuals of this diminutive whale were preyed on by the shark O. megalodon.
Charactosuchus
Charactosuchus is an extinct genus of crocodilian. It was assigned to the family Crocodylidae in 1988. Specimens have been found in Colombia, Brazil, Jamaica, and possibly Florida and South Carolina. It was gharial-like in appearance with its long narrow snout but bore no relation to them, being more closely related to modern crocodiles than to gharials.
Promacrauchenia
Promacrauchenia is an extinct genus of macraucheniids that lived during the Late Miocene to Late Pliocene epochs of what is now Argentina and Bolivia. It belongs to the subfamily Macraucheniinae, which also includes Huayqueriana, Macrauchenia, and Xenorhinotherium. Fossils of this genus have been found in the Ituzaingó, Andalhuala, and Cerro Azul Formations of Argentina.
Diplasiotherium
Diplasiotherium is an extinct genus of litoptern belonging to the family Proterotheriidae, that lived between the late Miocene and the early Pliocene (in the SALMAs Huayquerian and Montehermosan). The fossils of this animal have been found in Argentina, in the Monte Hermoso Formation.
Macroeuphractus
Macroeuphractus is a genus of extinct armadillos from the Late Miocene to Late Pliocene of South America. The genus is noted for its large size, with Macroeuphractus outesi being the largest non-pampathere or glyptodont armadillo discovered, as well as its specializations for carnivory, unique among all xenarthrans.
Megapaloelodus
Megapaloelodus is an extinct genus of stem flamingo of the family Palaelodidae. Megapaloelodus is primarily known from Miocene America, from South Dakota and Oregon in the north to Argentina in the south, but the species Megapaloelodus goliath was found in Europe. Additionally, one unnamed species was discovered in Miocene sediments from Namibia. Due to a lack of skull material, little can be said about the ecology of Megapaloelodus. Species of this genus are typically larger than those of Palaelodus and appear to have inhabited similar brackish lake environments. Additionally, they may have b
Macrauchenidia latidens
Huayqueriana is an extinct genus of South American litoptern, related to Macrauchenia, and belonging to the same family, Macraucheniidae. It was formerly known as Macrauchenidia latidens, described in 1939 by Cabrera, but redefined as Huayqueriana in 2016 based on the earlier name convention of Rovereto 1914. The genus is named after the Huayquerías Formation and the eponymous Huayquerian South American land mammal age defined at the formation.
Globidentosuchus
Globidentosuchus is an extinct genus of basal caimanine crocodylian known from the late Middle to Late Miocene of the Middle and the Upper Members of the Urumaco Formation at Urumaco, Venezuela. Its skull was very short and robust, with large units of spherical teeth used to break the shells of molluscs as part of its durophagus diet. It is thought to be one of the most basal Caimanines, even sharing some traits with alligatorids.
Paranauchenia denticulata
Paranauchenia is an extinct genus of South American litopterns belonging to the family Macraucheniidae. It is known only from fossil finds in Argentina. It possessed three toes and long limbs. The species Paranauchenia denticulata lived in the Miocene epoch in Argentina. Fossils have been found in the Arroyo Chasicó and Ituzaingó Formations of Argentina.
Solimoea acrensis
Solimoea acrensis is a prehistoric ateline monkey from the Late Miocene Solimões Formation of Brazil. It is the only known species of the genus Solimoea.
Pebas Formation
geological formation in Colombia