Middle Devonian first appearances

A tetrapod (; ) is any vertebrate animal of the clade Tetrapoda (). Tetrapods include all extant and extinct amphibians and amniotes, with the latter in turn evolving into two major clades, the sauropsids (reptiles, including dinosaurs and therefore birds) and synapsids (extinct "pelycosaurs", therapsids and all extant mammals, including humans).

Holocephali (sometimes spelled Holocephala; Greek for "complete head" in reference to the fusion of upper jaw with the skull) is a subclass of cartilaginous fish. The only living holocephalans are the three families of chimaeras, but the group also includes many extinct members and was more diverse during the Paleozoic and Mesozoic eras. The earliest known fossils of holocephalans date to the Middle Devonian, and the subclass likely reached its peak diversity during the following Carboniferous Period. Molecular clock studies suggest that holocephalans diverged from their closest relatives, ela

The Archaeognatha are an order of apterygotes, commonly known as bristletails. Among extant insect taxa they are some of the most evolutionarily primitive; they appeared in the Middle Devonian period at about the same time as the arachnids. Specimens that closely resemble extant species have been found as both body and trace fossils (the latter including body imprints and trackways) in strata from the remainder of the Paleozoic Era and more recent periods. For historical reasons an alternative name for the order is Microcoryphia.

thumb|right|Bark (possibly from a cladoxylopsid) from the Middle Devonian of Wisconsin

The progymnosperms are an extinct group of woody, spore-bearing plants that is presumed to have evolved from the trimerophytes, and eventually gave rise to the spermatophytes, ancestral to both gymnosperms and angiosperms (flowering plants). They have been treated formally at the rank of division Progymnospermophyta or class Progymnospermopsida (as opposite). The stratigraphically oldest known examples belong to the Middle Devonian order the Aneurophytales, with forms such as Protopteridium, in which the vegetative organs consisted of relatively loose clusters of axes. Tetraxylopteris is anoth

Goniatids, informally goniatites, are ammonoid cephalopods that form the order Goniatitida, derived from the more primitive Agoniatitida during the Middle Devonian some 390 million years ago (around Eifelian stage). Goniatites (goniatitids) survived the Late Devonian extinction to flourish during the Carboniferous and Permian only to become extinct at the end of the Permian some 139 million years later.

The Pituriaspida ('Pituri-shield' or 'hallucinogen-shield') are a small group of extinct armored jawless fishes with tremendous nose-like rostrums, which lived in the marine, deltaic environments of Middle Devonian Australia (about 393.3—382.7 Ma). They are known only by two species, Pituriaspis doylei and Neeyambaspis enigmatica found in a single sandstone location of the Georgina Basin, in Western Queensland, Australia.

Elpistostegalia is a clade containing Panderichthys and all more derived tetrapodomorph taxa. The earliest elpistostegalians, combining fishlike and tetrapod-like characters, such as Tiktaalik, are sometimes called fishapods. Although historically Elpistostegalia (referred to as Panderichthyida) was considered an order of prehistoric lobe-finned fishes, it was cladistically redefined to include tetrapods.

Tristichopterids (Tristichopteridae) were a diverse and successful group of fish-like tetrapodomorphs living throughout the Middle and Late Devonian. They first appeared in the Eifelian stage of the Middle Devonian. Within the group sizes ranged from a few tens of centimeters (Tristichopterus) to several meters (Hyneria and Eusthenodon).

Onychodus (, from Greek meaning "claw-tooth") is a genus of prehistoric lobe-finned fish which lived during the Devonian Period (Eifelian - Famennian stages, around 374 to 397 million years ago). It is one of the best known of the group of onychodontiform fishes. Scattered fossil teeth of Onychodus were first described from Ohio in 1857 by John Strong Newberry. Other species were found in Australia, England, Norway and Germany showing that it had a widespread range.

Heterosteus (also known as Heterostius) is an extinct genus of heterosteid placoderm of the Middle Devonian known from remains discovered in Europe and Greenland. According to Denison, 1978, Heterosteus might have been planktivorous, along with Homosteus, and Titanichthys.

The Archaeopteridaceae are an extinct family of plants belonging to Progymnospermae, and were dominant forest trees of the Late Devonian.

extinct genus of fishes

The Archaeopteridales are an extinct order of plants belonging to Progymnospermae, and dominant forest trees of the Late Devonian. They reproduced with spores rather than seeds. They were the evolutionary precursors of the conifers of gymnosperms which also include cycads and ginkgo.

Ceratarges is a genus of lichid Trilobite from the Middle to late Devonian. It lived in what is now western Europe and Morocco. It averaged around 10 cm (4 inches) in length and 9 cm (3.5 inches) in width.