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Santa Maria Formation

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Staurikosaurus
Staurikosaurus (Pronounced /ˌstɔɹ̠ikoʊˈsɔɹ̠ʌs/, STOR-ree-koh-SOR-ruhs; "Southern Cross lizard") is a genus of herrerasaurid dinosaur from the Late Triassic of Brazil, found in the Santa Maria Formation.
Saturnalia
extnct genus of dinosaurs
Hyperodapedon
Hyperodapedon (from , 'above' and , 'pavement') is an extinct genus of rhynchosaur reptiles which lived during Late Triassic period. Like other rhynchosaurs, it was a heavily built archosauromorph, distantly related to archosaurs such as crocodilians and dinosaurs. Hyperodapedon in particular was part of the subfamily Hyperodapedontinae, a specialized rhynchosaurian subgroup with broad skulls, beaked snouts, and crushing tooth plates on the roof of the mouth.
Dinodontosaurus
Dinodontosaurus (meaning "terrible-toothed lizard") is a genus of dicynodont therapsid. It was medium to large dicynodont of the Triassic (with skull up to long) and had a beak corneum. It lived in the Middle Triassic but disappeared in the Upper Triassic.
Buriolestes
Buriolestes is a genus of early sauropodomorph dinosaurs from the Late Triassic Santa Maria Formation of the Paraná Basin in southern Brazil. It contains a single species, B. schultzi, named in 2016. The type specimen was found alongside a specimen of the lagerpetid pterosauromorph Ixalerpeton.
Prestosuchus
Prestosuchus (meaning "Prestes crocodile") is an extinct genus of pseudosuchian in the group Loricata, which also includes Saurosuchus and Postosuchus. It has historically been referred to as a "rauisuchian", and was the defining member of the family Prestosuchidae, though the validity of both of these groups is questionable: Rauisuchia is now considered paraphyletic and Prestosuchidae is polyphyletic in its widest form.
Exaeretodon
Exaeretodon is an extinct genus of fairly large, low-slung traversodontid cynodonts from the southern parts of Pangea. Four species are known, hailing from various formations. E. argentinus is from the Carnian-age (Late Triassic) Cancha de Bochas Member of the Ischigualasto Formation in the Ischigualasto-Villa Unión Basin in northwestern Argentina. E. major and E. riograndensis are from the Carnian-age portion of the Santa Maria Formation of the Paraná Basin in southeastern Brazil. E. statisticae is from the Carnian-age Lower Maleri Formation of India.
Chiniquodon
thumb|left|Life reconstruction of C. theotonicus
Bagualosaurus
Bagualosaurus is a genus of sauropodomorph dinosaur from the Candelária Sequence (Uppermost Santa Maria Formation) of Brazil, dating to around 230 million years ago, in the Carnian of the Late Triassic. It includes one species, Bagualosaurus agudoensis.
Stahleckeria
Stahleckeria is an extinct genus of Middle Triassic (Ladinian) dicynodonts. It lived about 237 million years ago in what is now Brazil and Namibia. As a member of the group Kannemeyeriiformes, it was similar to the genus Kannemeyeria. The genus is known from the type species Stahleckeria potens, which was first collected from the Ladinian-age Santa Maria Formation in the Paleorrota fossil site of Brazil. Stahleckeria was named in honor of Rudolf Stahlecker, who discovered the first specimens during a 1935 expedition led by paleontologist Friedrich von Huene to the Chiniquá fossil site.
Rauisuchus
Rauisuchus (meaning "Wilhelm Rau's crocodile") is a genus of extinct archosaurs which lived in what is now the Geopark of Paleorrota (Santa María Formation), Brazil, during the Late Triassic period (235–228 million years ago). It contains one species, R. tiradentes.F. v. Huene. (1942) Die fossilen Reptilien des südamerikanischen Gondwanalandes. Ergebnisse der Sauriergrabung in Südbrasilien 1928/29. München: C.H. Beck'sche Verlagsbuchhandlung
Spondylosoma
Spondylosoma (meaning "vertebra body") is a genus of avemetatarsalian archosaur, belonging to the clade Aphanosauria, from the Middle Triassic (late Ladinian age) Lower Santa Maria Formation in Paleorrota Geopark, Brazil.
Pampadromaeus
Pampadromaeus is an extinct genus of basal sauropodomorph dinosaurs known from the Late Triassic (Carnian) Santa Maria Formation of the Paraná Basin in Rio Grande do Sul, southern Brazil.
Gnathovorax
Gnathovorax is a genus of herrerasaurid saurischian dinosaur from the Santa Maria Formation in Rio Grande do Sul, Brazil. The type and only species is Gnathovorax cabreirai, described by Pacheco et al. in 2019. The type specimen comprises a mostly complete and well-preserved articulated skeleton (considered one of the best herrerasaurid skeletons ever discovered), which fossilized in close association with rhynchosaur and cynodont remains. The discovery of this superb specimen has shed light onto poorly understood aspects of herrerasaurid anatomy such as endocranial soft tissues.
Massetognathus
Massetognathus ( ; Greek for "chewing muscle jaw") is an extinct genus of plant-eating traversodontid cynodonts. They lived during the Triassic Period about 235 million years ago, and are known from the Chañares Formation in Argentina and the Santa Maria Formation in Brazil.
Trucidocynodon
Trucidocynodon is an extinct genus of ecteniniid cynodonts from the Upper Triassic (Carnian) of Brazil. It is a monotypic genus that contains a single species, Trucidocynodon riograndensis. Fossils of Trucidocynodon were discovered in outcrops of the Upper Santa Maria Formation in Paleorrota Geopark, Agudo. Trucidocynodon is one of the most completely known Triassic cynodonts, as its holotype is a nearly complete and fully articulated skeleton.
Erythrovenator
Erythrovenator is a genus of basal theropod dinosaurs from the Late Triassic of Rio Grande so Sul, Brazil. The genus contains a single species, Erythrovenator jacuiensis.
Ixalerpeton
Ixalerpeton (meaning "leaping reptile") is a genus of lagerpetid avemetatarsalian containing one species, I. polesinensis. It lived in the Late Triassic of Brazil alongside the sauropodomorph dinosaur Buriolestes.
Nhandumirim
Nhandumirim (meaning "small rhea" in the Tupi language) is a genus of basal sauropodomorph dinosaur from the Carnian age of Late Triassic Brazil. It is currently considered a saturnaliid sauropodomorph. The type and only species, Nhandumirim waldsangae, is known from a single immature specimen including vertebrae, a , pelvic material, and a hindlimb found in the Santa Maria Formation in Rio Grande do Sul.
Bonacynodon
Bonacynodon is an extinct genus of cynodonts that lived in what is now southern Brazil during the Triassic period (Ladinian–Carnian ages). The genus is monotypic, containing only the type species Bonacynodon schultzi. B. schultzi is known from two specimens, consisting of two partial skulls and some badly preserved parts of the postcranium. Both specimens were recovered from the Pinheiros-Chiniquá Sequence, part of the Santa Maria Supersequence of the Paraná Basin. This sequence preserves a faunal association known as the Dinodontosaurus Assemblage Zone, which contains numerous other species o
Proterochampsa
Proterochampsa is an extinct genus of proterochampsid archosauriform from the Late Triassic (Carnian-Norian boundary) of South America. The genus is the namesake of the family Proterochampsidae, and the broader clade Proterochampsia. Like other proterochampsids, Proterochampsa are quadruped tetrapods superficially similar in appearance to modern crocodiles, although the two groups are not closely related. Proterochampsids can be distinguished from other related archosauriformes by characters such as a dorsoventrally flattened, triangular skull with a long, narrow snout at the anterior end and
Chanaresuchus
Chanaresuchus is an extinct genus of proterochampsid archosauriform. It was of modest size for a proterochampsian, being on average just over a meter in length. The type species is Chanaresuchus bonapartei was named in 1971. Its fossils were found in from the early Carnian-age Chañares Formation in La Rioja Province, Argentina . Chanaresuchus appears to be one of the most common archosauriforms from the Chañares Formation due to the abundance of specimens referred to the genus. Much of the material has been found by the La Plata-Harvard expedition of 1964–65. Chanaresuchus is the most well-des
Candelariodon
Candelariodon is an extinct genus of carnivorous probainognathian cynodonts from the Middle to Late Triassic Santa Maria Formation of the Paraná Basin in Rio Grande do Sul state, Brazil. Candelariodon is known from a partial mandible having some complete teeth. It was first named by Téo Veiga de Oliveira, Cesar Leandro Schultz, Marina Bento Soares and Carlos Nunes Rodrigues in 2011 and the type species is Candelariodon barberenai.
Decuriasuchus quartacolonia
Decuriasuchus is an extinct genus of loricatan from the Middle Triassic period (Ladinian stage). It is a carnivorous archosaur that lived in what is now southern Brazil, in Paleorrota. It was first named by Marco Aurélio G. França, Jorge Ferigolo and Max C. Langer in 2011 and the type species is Decuriasuchus quartacolonia. The generic name means "unit of ten crocodiles" in Latin and Greek in reference to the ten known specimens and the animal's possible group behavior. The specific name refers to the Quarta Colonia region where the fossils were collected.
Protuberum
Protuberum is an extinct genus of traversodontid cynodonts known from a single species Protuberum cabralense, from the Middle Triassic of Brazil.
Ginkgoites
Ginkgoites is a genus of extinct plants belonging to Ginkgoaceae. Fossils of these plants have been found around the globe during the Triassic, Jurassic, Cretaceous, with fossils also known from the Paleogene. The name was created as a form genus in 1919 by Albert Seward, who stated: "I... propose to employ the name Ginkgoites for leaves that it is believed belong either to plants generically identical with Ginkgo or to very closely allied types."
Menadon
Menadon is an extinct genus of traversodontid cynodonts. The type and only species is Menadon besairiei.
Archeopelta
Archeopelta is an extinct genus of carnivorous archosaur from the late Middle or early Late Triassic period (late Ladinian to early Carnian stage). It was a 2 m (6 ft) long predator which lived in what is now southern Brazil. Its exact phylogenetic placement within Archosauriformes is uncertain; it was originally classified as a doswelliid, but subsequently it was argued to be an erpetosuchid archosaur.
Santa Maria Formation
formation
Venetoraptor
Venetoraptor (meaning "Vale Vêneto plunderer") is an extinct genus of lagerpetid archosaurs from the Late Triassic Santa Maria Formation (Paraná Basin) of Rio Grande do Sul, Brazil. The genus contains a single species, V. gassenae, known from a partial skeleton. Venetoraptor is characterized by the presence of a beak and scythe-like claws on long fingers.
Polesinesuchus
Polesinesuchus is an extinct genus of stagonolepidid aetosaur known from the Late Triassic of southern Brazil. Fossils have been found from the Santa Maria Supersequence of the late Carnian and early Norian stages, making Polesinesuchus one of the oldest aetosaurs. It contains a single species, Polesinesuchus aurelioi, the fifth aetosaur species known from South America to date. Anatomical evidence suggests that Polesinesuchus likely represents a juvenile individual of the contemporary Aetosauroides.
Rhadinosuchus
Rhadinosuchus is an extinct genus of proterochampsian archosauriform reptile from the Late Triassic. It is known only from the type species Rhadinosuchus gracilis, reposited in Munich, Germany. The fossil includes an incomplete skull and fragments of post-cranial material. Hosffstetter (1955), Kuhn (1966), Reig (1970) and Bonaparte (1971) hypothesized it to be synonymous with Cerritosaurus, but other characteristics suggest it is closer to Chanaresuchus and Gualosuchus, while it is certainly different from Proterochampsa and Barberenachampsa. The small size indicates it is a young animal, maki
Aetobarbakinoides
Aetobarbakinoides is an extinct genus of stagonolepidid aetosaur known from the Late Triassic of Rio Grande do Sul state, southern Brazil. Fossils have been found from the Santa Maria Supersequence of the late Carnian and early Norian stages, making Aetobarbakinoides one of the oldest aetosaurs. The type species, A. brasiliensis, was named in 2012, and is notable for being described primarily by features of the vertebrae; most other aetosaurs are diagnosed by features in bony plates called osteoderms, which are by far the most common material. Although placed in a basal phylogenetic position a
Cerritosaurus
Cerritosaurus is a genus of proterochampsid archosauromorph from the Late Triassic. It has been found in the Santa Maria Formation, in the Geopark of Paleorrota, Brazil. It is represented by one species.
Therioherpeton
Therioherpeton is an extinct genus of small, carnivorous cynodonts belonging to the clade Prozostrodontia, which lived in what is now Brazil during the Late Triassic. Its type species is Therioherpeton cargnini. It was named in 1975 by the palaeontologists José Bonaparte and Mário Costa Barberena based on remains collected in the Hyperodapedon Assemblage Zone of the Santa Maria Formation in the Paraná Basin.
Aetosauroides
Aetosauroides (meaning "Aetosaurus-like") is an extinct genus of aetosaur from the Late Triassic of South America. It is one of four aetosaurs known from South America, the others being Neoaetosauroides, Chilenosuchus and Aetobarbakinoides. Three species have been named: the type species A. scagliai, A. subsulcatus, and A. inhamandensis. Fossils have been found in the Cancha de Bochas Member of the Ischigualasto Formation in the Ischigualasto-Villa Unión Basin in northwestern Argentina and the Santa Maria Formation in the Paraná Basin in southeastern Brazil. The strata date to the late Carnian
Charruodon
Charruodon is an extinct genus of cynodonts which existed in the Hyperodapedon Assemblage Zone of the Santa Maria Formation in the Paraná Basin in southeastern Brazil during the Late Triassic. The genus contains only the type species Charruodon tetracuspidatus, which is known from a single specimen of uncertain provenance. Upon its first description, Charruodon was tentatively placed within the family Therioherpetidae, but a 2017 study by Agustín G. Martinelli and colleagues instead recovered it as a more basal member of Probainognathia. In 2023, Hoffmann, Ribeiro & de Andrade reinterpreted th
Protheriodon
Protheriodon is an extinct genus of probainognathian cynodonts which existed in the Santa Maria Formation of the Paraná Basin in southeastern Brazil during the middle Triassic period. It contains the species Protheriodon estudianti. It was first described by Argentine palaeontologist José Bonaparte, who assigned it to the family Brasilodontidae. More recent studies have however recovered it in a more basal position than other brasilodontids, just outside Prozostrodontia.
Parvosuchus
Parvosuchus (meaning "small crocodile") is an extinct genus of gracilisuchid pseudosuchians from the Middle–Late Triassic Santa Maria Formation (Paraná Basin) of Brazil. The genus contains a single species, P. aurelioi, known from a cranium and partial skeleton.
Dagasuchus
Dagasuchus is an extinct genus of pseudosuchian archosaur from the Late Triassic (Carnian) of Rio Grande do Sul, Brazil, represented by the type species Dagasuchus santacruzensis. D. santacruzensis was named in 2015 on the basis of specimen UFRGS-PV-1244-T and UFRGS-PV-1245-T, a partial hip (one ilium and a pair of ischia) found in an exposure of the Santa Maria Formation in the Paraná Basin, near the city of Santa Cruz do Sul. Dagasuchus is an early member of a large evolutionary group called Loricata, which originated in the Triassic and includes modern crocodylians and their ancestors. Feat
Candelaria
species of reptile (fossil)
Santacruzgnathus
Santacruzgnathus is an extinct genus of small cynodonts from the Late Triassic (Carnian) Santacruzodon Assemblage Zone of Brazil. It contains one species, S. abdalai. Santacruzgnathus is known from a single partial lower jaw with four postcanine teeth, only one of which is well-preserved. Some features of the specimen, including the slender shape of the jaw and the incipiently double-rooted teeth, indicate that the animal was an early member of Prozostrodontia, a group that includes mammals and their close relatives.
Alemoatherium
Alemoatherium is an extinct genus of prozostrodontian cynodont which lived in the Late Triassic of Brazil. It contains a single species, A. huebneri, named in 2017 by Agustín Martinelli and colleagues. The genus is based on UFSM 11579b, a left lower jaw (dentary) found in the Alemoa Member of the Santa Maria Formation, preserving the late Carnian-age Hyperodapedon Assemblage Zone. Alemoatherium was among the smallest species of cynodonts found in the rich synapsid fauna of the Santa Maria Formation. Its blade-like four-cusped postcanine teeth show many similarities with those of dromatheriids,