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Wheeler Shale

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Anomalocaris
Anomalocaris (from Ancient Greek , meaning "unlike", and , meaning "shrimp", with the intended meaning "unlike other shrimp") is an extinct genus of radiodont, an order of early-diverging stem-group marine arthropods. It is best known from the type species A. canadensis, found in the Stephen Formation (particularly the Burgess Shale) of British Columbia, Canada. The other named species A. daleyae is known from the somewhat older Emu Bay Shale of Australia. Other unnamed Anomalocaris species are known from China and the United States.
Hallucigenia
Hallucigenia is a genus of lobopodian known from Cambrian-aged fossils in Burgess Shale-type deposits in Canada (Burgess Shale) and China, and from isolated spines around the world. The generic name reflects the type species' unusual appearance and eccentric history of study; when it was erected as a genus, H. sparsa was reconstructed as an enigmatic animal upside down and back to front. Lobopodians are a grade of Paleozoic panarthropods from which the velvet worms, water bears, and arthropods arose.
Wiwaxia
Wiwaxia is a genus of prehistoric soft-bodied animals that were covered in carbonaceous scales and spines that protected it from predators. Wiwaxia fossils—mainly isolated scales, but sometimes complete, articulated fossils—are known from early Cambrian and middle Cambrian fossil deposits across the globe. The living animal would have measured up to when fully grown, although a range of juvenile specimens are known, the smallest being long.
Aysheaia
Aysheaia is an extinct genus of soft-bodied lobopodian, known from the Middle Cambrian Burgess Shale of British Columbia, Canada.
Ottoia
Ottoia is a stem-group archaeopriapulid worm known from Cambrian fossils. Although priapulid-like worms from various Cambrian deposits are often referred to Ottoia on spurious grounds, the only clear Ottoia macrofossils come from the Burgess Shale of British Columbia, which was deposited . Microfossils extend the record of Ottoia throughout the Western Canada Sedimentary Basin, from the mid- to late Cambrian. A few fossil finds are also known from China.
Alalcomenaeus
Alalcomenaeus is one of the most widespread and longest-surviving arthropod genera of the Early and Middle Cambrian. Known from over 300 specimens in the Burgess Shale and the Chengjiang biota. It is a member of the family Leanchoiliidae in the group Megacheira.
Naraoia
Naraoia is a genus of small-to-average-size (about long) marine arthropods within the family Naraoiidae, that lived from the early Cambrian to the late Silurian period. The species are characterized by a large alimentary system and sideways oriented antennas.
Sidneyia
Sidneyia is an extinct marine arthropod known from fossils found from the Early to the Mid Cambrian of China and the Mid Cambrian Burgess Shale of British Columbia, Canada. It is thought to have been a durophagous (crushing) predator that used its spine-covered leg bases (gnathobases) to feed on hard-bodied animals like trilobites and brachiopods, as well as possibly soft bodied animals. It was historically placed in the group "Xenopoda" alongside Emeraldella. It is now placed within the clade Vicissicaudata along with aglaspidids, chelonellids, as well as Emeraldella, among others. Vicissicau
Canadaspis
Canadaspis ("Shield of Canada") is an extinct genus of bivalved Cambrian marine arthropod, known from North America and China. They are thought to have been benthic feeders that moved mainly by walking and possibly used its biramous appendages to stir mud in search of food. They have been placed within the Hymenocarina, which includes other bivalved Cambrian arthropods.
Leanchoilia
Leanchoilia is a megacheiran marine arthropod known from Cambrian deposits of the Burgess Shale in Canada and the Chengjiang biota of China.
Waptia
Waptia is an extinct genus of marine arthropod from the Middle Cambrian of North America. It grew to a length of , and had a large bivalved carapace and a segmented body terminating into a pair of tail flaps. It was an active swimmer and likely a predator of soft-bodied prey. It is also one of the oldest animals with direct evidence of brood care. Waptia fieldensis is the only species classified under the genus Waptia, and is known from the Burgess Shale Lagerstätte of British Columbia, Canada. Specimens of Waptia are also known from the Spence Shale of Utah, United States.
Eldonia
Eldonia is an extinct soft-bodied cambroernid best known from the Fossil Ridge outcrops of the Burgess Shale, particularly in the 'Great Eldonia layer' in the Walcott Quarry. In addition to over 550 specimens collected by Walcott, 224 specimens of Eldonia are known from the Greater Phyllopod bed, where they comprise 0.43% of the community. Species also occur in the Chengjiang biota, and Siberia.
Olenoides
Olenoides is a genus of trilobite from the Cambrian period. Its fossils can be found with soft body parts intact in the Burgess Shale in Canada. Fossils of this genus can also be found in the Wheeler Shale and Marjum Formation of Utah in the United States of America, among other localities. Species within this genus range in size, with most being medium-sized trilobites (Olenoides Serratus from the Burgess Shale can be up to 9 centimeters in length). Specimens of Olenoides Superbus, pictured on the right, an especially large species, can reach sizes of well over 10 centimeters in length. Oleno
Elrathia
thumb|right|250px|Elrathia kingii growth series with holaspids ranging from 16.2 mm to 39.8 mm in length
Perspicaris
Perspicaris (from the Latin perspicax, meaning “sharp-sighted,” and caris, “crab/shrimp”) an extinct genus of bivalved arthropod from the Cambrian period. Fossils have been found in North America, primarily the Burgess Shale of British Columbia, Canada but also possibly the Wheeler Shale, Marjum Formation, Pioche Shale and Bloomington Formation. Two named species are known from the Burgess Shale Perspicaris dictynna and Perspicaris recondita, which differ in maximum size ( in P. recondita vs in P. dictynna), as well as proportions of the tail. Both species have a pair of stalked eyes, as well
Tuzoia
Tuzoia (from Mount Tuzo, a mountain in the Canadian Rockies) is an extinct genus of large bivalved arthropod known from Early to Middle Cambrian marine environments from what is now North America, Australia, China, Europe and Siberia. The large, domed carapace reached lengths of , making them amongst the largest known Cambrian arthropods.
Isoxys
Isoxys (meaning "equal surfaces") is a genus of extinct bivalved Cambrian arthropod; the various species of which are thought to have been freely swimming predators. It had a pair of large spherical eyes (which are the most commonly preserved feature of the soft-bodied anatomy), and two large frontal appendages used to grasp prey.
Choia
Choia is a genus of extinct demosponge ranging from the Cambrian until the Lower Ordovician periods. Fossils of Choia have been found in the Burgess Shale in British Columbia; the Maotianshan shales of China; the Wheeler Shale in Utah; the Itajaí Basin in Brazil; and the Lower Ordovician Fezouata formation. It was first described in 1920 by Charles Doolittle Walcott.
Asaphiscus
Asaphiscus is a genus of trilobite that lived in the Cambrian. Its remains have been found in Australia and North America, especially in Utah.
Cothurnocystis
Cothurnocystis is a genus of small enigmatic echinoderms that lived during the Ordovician. Individual animals had a flat boot-shaped body and a thin rod-shaped appendage that may be a stem, or analogous to a foot or a tail. Fossils of Cothurnocystis species have been found in Nevada, Scotland, Czech Republic, France and Morocco.
Kootenia
thumb|left|Kootenia assemblage at the Geological Museum in Copenhagen
Selkirkia
Selkirkia is a genus of predatory, tubicolous priapulid worms known from the Middle Cambrian Burgess Shale, Ogygopsis Shale, Puncoviscana Formation and the Early Ordovician Fezouata Formation. 142 specimens of Selkirkia are known from the Greater Phyllopod bed, where they comprise 0.27% of the community. In the Burgess Shale, 20% of the tapering, organic-walled tubes are preserved with the worm inside them, whereas the other 80% are empty (or sometimes occupied by one or more small agnostid trilobites). Whilst alive, the tubes were probably vertical, whereas trilobite-occupied tubes are horizo
Branchiocaris
Branchiocaris is an extinct genus of Cambrian bivalved arthropod. The type and best known species, Branchiocaris pretiosa, was described from the Burgess Shale of British Columbia, Canada, in 1929, originally placed in Protocaris, and was placed into its own distinct genus by Briggs in 1976. Several other possible species have been described from Cambrian deposits in China, and it is also possibly known from Cambrian deposits in Utah.
Skeemella
Skeemella is a genus of elongate animal from the Middle Cambrian Wheeler Shale and Marjum lagerstätte of Utah. It has been classified with the banffozoan vetulicolians.
Emeraldella
Emeraldella is a genus of arthropod known from the Middle Cambrian of North America. The type species E. brocki was described in 1912 from the Burgess Shale. 21 specimens of Emeraldella are known from the Greater Phyllopod bed, where they comprise < 0.1% of the community. A re-study on the species was done in 2012. A second species E. brutoni is known from the Wheeler Shale, which was described in 2011. An additional specimen of E. brutoni was described in 2019, which revealed more of the anatomy. It has been placed as a basal member of the clade Vicissicaudata within Artiopoda, a group of art