Calomela

Length: 4 - 11 mm; body shape variable, most species moderately narrow (length to width ratio 1.8:1) and flat (length to height ratio 2.6:1), but some (generally rainforest species) broad (length to width ratio < 1.8:1) and convex (length to height ratio ranges from 2:1 to 3.3:1). Head: Contracted behind eyes; frons without vertical groove beside inner margin of eye; frontoclypeal suture strongly transverse, V shaped or rounded; first segment of maxillary palp ventrally convex, without sharp leading edge; apical segment of maxillary palp strongly expanded from base to truncate apex; apical margin of mentum truncate or shallowly concave. Thorax: Pronotum broadest at middle or at base; anterior trichobothrium present ; posterior trichobothrium present; sides of pronotal disc with irregular punctate depression on each side, rarely with only scattered large punctures (eg C. jansoni ); base of pronotum without margination; hypomeral groove present, curving from prosternal suture then parallel to pronotal margin, rarely shallow ( C. relicta ); anterior of prosternum without median or lateral ridges, midline not anteriorly produced; prosternal process narrow, longer than broad (but apex often bilobed or abruptly expanded), with basal right-angled lobes absent; procoxal cavity open, gap at least half width of procoxa; elytral tubercules absent; elytral striae usually present, rarely absent (but punctures longitudinally arranged), 5th stria deeper at base compared to adjacent striae, or not; elytra not extended vertically, epipleura entirely visible; base of elytra not overlapping basal angles of pronotum; epipleuron narrow, < 0.2 times width of elytron, without setae ; anterior face of mesoventrite process rarely gradually raised, usually abruptly raised and convex or truncate, posterior margin truncate to strongly concave; metaventrite anterior femoral plates absent; metaventrite process not raised anteriorly; apices of mid and hind tibiae without row of short spines on distal surface; tibiae without obvious external keels; apex of third tarsal segment not or feebly bilobed; claws simple, or simply toothed or appendiculate with sharp basal tooth. Abdomen: Pygidial groove absent; abdominal ventrites with or without patches of large setiferous punctures; apex of last ventrite strongly or shallowly excavate, or truncate, often with a pair of teeth, usually with median depression in male. The genus Calomela occurs throughout Australia and is also present in New Guinea. In Australia there are about 45 species, at least 20 occurring in New South Wales (Selman 1976, 1977, 1979; Reid 1989, 1992). Most species are common where they occur. Calomela species are particularly associated with Acacia and more than one species may occur on a single plant. For example, a small plantation of Acacia mearnsii near Queanbeyan included co-occuring populations of four species of Calomela (Hunt et. al. 1996). In most species, adults and larvae are leaf feeding but some adults prefer flowers. Host specificity appears to be rare, most Calomela species with recorded hosts have more than one host species (Reid 1989, 1992). The concept of this genus has changed in recent years to include several other poorly distinguished genera (Daccordi 1994; Reid 2004). The species have similar biology where known.