Cimoliopterus

Cimoliopterus is a genus of pterosaur that lived during the Late Cretaceous in what is now England and the United States. The first known specimen, consisting of the front part of a snout including part of a crest, was discovered in the Grey Chalk Subgroup of Kent, England, and described as the new species Pterodactylus cuvieri in 1851. The specific name cuvieri honoured the palaeontologist George Cuvier, and the genus Pterodactylus was then used for many pterosaurs of species that are no longer thought to be closely related. It was among the first pterosaurs to be depicted as sculptures, in Crystal Palace Park in the 1850s. The species was subsequently assigned to various other genera, including Ornithocheirus and Anhanguera. In 2013, the species was moved to a new genus, as C. cuvieri; the generic name Cimoliopterus is derived from the Greek words for "chalk" and "wing". Other specimens and species have also been assigned to or synonymised with the species with various levels of certainty. In 2015, a snout discovered in the Britton Formation of Texas, US, was named as a new species in the genus, C. dunni. A species from the Cambridge Greensand of England originally assigned to Ornithocheirus was assigned to Cimoliopterus in 2025, as C. colorhinus.

C. cuvieri is estimated to have had a wingspan of , and C. dunni is thought to have been similar to C. cuvieri in size. One C. colorhinus specimen is estimated to have had a wingspan of . Cimoliopterus can be distinguished from related pterosaurs in features such as having a premaxillary crest that begins hindward on the snout, in having a ridge on the palate that extends forwards until the third pair of tooth sockets, and in the spacing and proportions of the tooth sockets. Unlike similar pterosaurs, the tip of the snout is only subtly expanded to the sides. C. cuvieri and C. dunni differ from each other in various details in the configuration of these features; for example, the crest of C. cuvieri begins by the seventh tooth socket, whereas that of C. dunni begins at the fourth. More completely known related genera were fairly large pterosaurs, with proportionally large skulls, long jaws and tooth-rows, often with large, rounded crests at the front of the jaws. The teeth at the front of the jaws were large and recurved; further back, the teeth were smaller, slightly recurved, and well-spaced. As pterosaurs, Cimoliopterus would have been covered in pycnofibres (hair-like filaments), and had extensive wing-membranes, which were distended by long wing-fingers.