Cladoselache

Cladoselache ("branch shark") is an extinct genus of shark-like chondrichthyan (cartilaginous fish) from the Late Devonian (Famennian) of North America. It was similar in body shape to modern lamnid sharks (such as mako sharks and the great white shark), but was not closely related to lamnids or to any other modern (selachian) shark. As an early chondrichthyan, it had yet to evolve traits of modern sharks such as accelerated tooth replacement, a loose jaw suspension, enameloid teeth, and possibly claspers.

Some 20th century studies considered Cladoselache to be a basal (early-diverging) member of Elasmobranchii, the fork of cartilaginous fish which leads to modern sharks and rays. More recent studies have identified distinctive traits of the chondrocranium (cartilaginous braincase), dorsal fin spines, and pectoral fin bases. These newly identified features support a close relationship to symmoriiforms, a small group of bizarre chondrichthyans such as the bristle-spined Stethacanthus. Cladoselache and symmoriiforms may be more closely related to chimaeras (a modern group of unusual deep-sea fish) than to true sharks and rays.