Gorgonopsia

Gorgonopsia (from the Greek Gorgon, a mythological beast, and 'aspect') is an extinct clade of sabre-toothed therapsids from the Middle to the Upper Permian, roughly between 270 and 252 million years ago. They are characterised by a long and narrow skull, as well as elongated upper and sometimes lower canine teeth and incisors which were likely used as slashing and stabbing weapons. Postcanine teeth are generally reduced or absent. For hunting large prey, they possibly used a bite-and-retreat tactic, ambushing and taking a debilitating bite out of the target, and following it at a safe distance before its injuries exhausted it, whereupon the gorgonopsian would grapple the animal and deliver a killing bite. They would have had an exorbitant gape, possibly in excess of 90°, without having to unhinge the jaw.

They markedly increased in size as time went on, growing from small skull lengths of in the Middle Permian to bear-like proportions of up to in the Upper Permian. The latest gorgonopsians, Rubidgeinae, were the most robust of the group and could produce especially powerful bites. Gorgonopsians are thought to have been completely terrestrial and could walk with a semi-erect gait, with a similar terrestrial locomotory range as modern crocodilians. They may have been more agile than their prey items, but were probably inertial homeotherms rather than endotherms unlike contemporary therocephalians and cynodonts, and thus were probably comparatively less active. Though gorgonopsians were able to maintain a rather high body temperature, it is unclear if they would have also had sweat glands or fur (and by extension whiskers and related structures). Their brains were reminiscent of modern reptilian brains, rather than those of living mammals. Most species may have been predominantly diurnal (active during the day) though some could have been crepuscular (active at dawn or dusk) or nocturnal (active at night). They are thought to have had binocular vision, a parietal eye (which detects sunlight and maintains circadian rhythm), a keen sense of smell, a functional vomeronasal organ ("Jacobson's organ"), and possibly a rudimentary eardrum.