Leptothecata

Leptothecata, or thecate hydroids, are an order of hydrozoans in the phylum Cnidaria. Their closest living relatives are the athecate hydroids, which are similar enough to have always been considered closely related, and the very apomorphic Siphonophorae, which were placed outside the "Hydroida". Given that there are no firm rules for synonymy for high-ranked taxa, alternative names like Leptomedusa, Thecaphora or Thecata, with or without the ending emended to "-ae", are also often used for Leptothecata.

In the sessile stage, Leptothecata are surrounded by a chitinous outer layer as their exoskeleton, including the gonophores, their reproductive organ. Leptothecata exhibit radial symmetry, and their gonads can be found in the radial canals of the medusa stage. Their habits range from benthic to planktonic. The polyps and colonial forms are benthic, whilst the medusae are planktonic. Leptothecata exhibit extensive, complex variation. Thecata colonies also have extensive specialization due to their polyps' function and variation. Most Leptothecata possess statocysts, which are used for defence and protection. The classes that have lost their statocysts have been changed ancestrally over time rather than a direct loss.