Sanctacaris

Sanctacaris is a Middle Cambrian arthropod from the Burgess Shale of British Columbia. It was most famously regarded as a stem-group chelicerate, a group which includes horseshoe crabs, spiders and scorpions, although subsequent phylogenetic studies have not always supported this conclusion. Its chelicerate affinities regain support in later observations, alongside the reassignment of Habelia optata as a sanctacaridid-related basal chelicerate. It has been placed as a member of the extinct family Sanctacarididae alongside Wisangocaris and Utahcaris.

Sanctacaris specimens range from 46 to 93 mm in length. The head bears five pairs of grasping appendages (corresponding to chelicerate's pedipalps and walking legs) and a sixth pair of posterior appendages (correspond to horseshoe crab's chilaria). The grasping appendages each bear an antenna-like exopods. There are 11 body segments, with the former 10 each bearing a pair of biramous appendages with flap-like exopod and reduced leg-like endopod. There is a broad, flat paddle-like telson.