Thalassodromeus

Thalassodromeus is a genus of pterosaur that lived in what is now Brazil during the Early Cretaceous period, about a hundred million years ago. The original skull, discovered in 1983 in the Araripe Basin of northeastern Brazil, was collected in several pieces. In 2002, the skull was made the holotype specimen of Thalassodromeus sethi by palaeontologists Alexander Kellner and Diogenes de Almeida Campos. The generic name means "sea runner" (in reference to its supposed mode of feeding), and the specific name refers to the Egyptian god Seth due to its crest being supposedly reminiscent of Seth's crown. Other scholars have pointed out that the crest was instead similar to the crown of Amon. A jaw tip was assigned to T. sethi in 2005, became the basis of the new genus Banguela in 2015, and assigned back to Thalassodromeus as the species T. oberlii in 2018, though other researchers consider it a valid genus. Another species (T. sebesensis) was described in 2015 based on a supposed crest fragment, but this was later shown to be part of a turtle shell.

Thalassodromeus had one of the largest known skulls among pterosaurs, around long, with one of the proportionally largest cranial crests of any vertebrate. Though only the skull is known, the animal is estimated to have had a wingspan of . The crest was lightly built and ran from the tip of the upper jaw to beyond the back of the skull, ending in a unique V-shaped notch. The jaws were toothless, and had sharp upper and lower edges. Its skull had large nasoantorbital fenestrae (opening that combined the antorbital fenestra in front of the eye with the bony nostril), and part of its palate was concave. The lower jaw was blade-like, and may have turned slightly upwards. The closest relative of Thalassodromeus was Tupuxuara; both are grouped in a clade called Thalassodromidae, which, depending on the study, has been placed either within Tapejaromorpha, closely related to the family Tapejaridae, or within Neoazhdarchia, closely related to the family Dsungaripteridae.