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Graphidales genera

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Phaeographis
Phaeographis is a genus of lichen-forming fungi in the family Graphidaceae. It has an estimated 180 species.
Graphis
genus of fungi
Diploschistes
Diploschistes is a genus of lichen-forming fungi in the family Graphidaceae. Commonly known as crater lichens, members of the genus are crustose lichens with a thick, cracked (areolate) body (thallus) with worldwide distribution. The fruiting part (apothecia) are immersed in the thick thallus so as to have the appearance of being small "craters". The widespread genus contains about 43 species.
Gyalidea
Gyalidea is a genus of crustose lichens in the family Gomphillaceae. It has 50 species.
Thelotrema
Thelotrema is a genus of lichen-forming fungi in the family Graphidaceae, the family to which all taxa in the former Thelotremataceae now belong.
Gyalectidium
Gyalectidium is a genus of lichen-forming fungi in the family Gomphillaceae. Established in 1881 by Swiss lichenologist Johannes Müller Argoviensis, the genus comprises about 50 species of predominantly leaf-dwelling lichens that form small, pale greenish to whitish-grey patches typically only a few millimetres across. Members are characterised by specialised asexual reproductive structures called , which are small scale-like outgrowths that produce propagules containing both fungal and algal partners, and by fruiting bodies that contain single, large, multi-chambered ascospores. The genus has
Gymnographa
Gymnographa is a genus of lichen-forming fungi in the family Graphidaceae. It contains two species.
Jamesiella
Jamesiella is a genus of lichen-forming fungi in the family Gomphillaceae. Members of Jamesiella form thin, delicate crusts on tree bark, rocks, and mosses in humid tropical and temperate forests, distributed across North and South America and Europe. The genus is distinguished from its close relative Gyalideopsis by a unique type of asexual reproductive structure called , which are specialized stalks containing both fungal filaments and algal cells that detach and disperse as complete units capable of establishing new lichens.
Asterothyrium
Asterothyrium is a genus of leaf-dwelling lichens in the family Gomphillaceae. These tiny lichens form small, star-shaped fruiting bodies on the surface of leaves in tropical and subtropical forests. They are most commonly found in humid, shaded environments where leaves remain moist and clean. The genus includes 19 recognised species distributed primarily throughout Central and South America.
Gyalideopsis
Gyalideopsis is a genus of lichen-forming fungi in the family Gomphillaceae. Gyalideopsis species form delicate, film-like crusts on a wide variety of surfaces including tree bark, rocks, and mosses, often in tropical and temperate forests worldwide. Species in the genus produce distinctive small, scale-like structures called , which are thought to aid in asexual reproduction where ascospore-producing structures are absent or uncommon. Members of this genus lack the distinctive chemical compounds found in many other lichens, making them reliant on microscopic features and spore characteristics
Phaeographina
Phaeographina is a genus of lichen-forming fungi in the family Graphidaceae. The genus was circumscribed by Swiss lichenologist Johannes Müller Argoviensis in 1882.
Echinoplaca
Echinoplaca is a genus of lichen-forming fungi in the family Gomphillaceae.
Tricharia
Tricharia is a genus of lichens in the family Gomphillaceae. It has an estimated 30 species.
Carbacanthographis
Carbacanthographis is a genus of corticolous (bark-dwelling) lichens in the family Graphidaceae. The genus was circumscribed by the German lichenologists Bettina Staiger and Klaus Kalb in 2002. An updated worldwide key to the genus was published in 2022 that added 17 new species. This revision allowed for further identification of undescribed species from other collections, and subsequently, 14 species were added in 2023 from the Amazonian lowland forests of Brazil and the Guianas.
Sarcographa
Sarcographa is a genus of lichen-forming fungi in the family Graphidaceae. Established in 1825 by the French botanist Antoine Laurent Apollinaire Fée, the genus contains 22 species that are recognised by their distinctive star-shaped colonies of radiating, script-like fruiting structures with black borders. These bark-dwelling lichens are found in humid tropical and warm temperate forests worldwide and serve as indicators of undisturbed woodland, as they quickly decline when forest canopy is opened or disturbed.
Psorotheciopsis
Psorotheciopsis is a genus of lichenized fungi in the family Gomphillaceae. It contains three species.
Acanthothecis
Acanthothecis is a genus of lichen-forming fungi in the family Graphidaceae. The genus was circumscribed by Frederick Edward Clements in 1909. These lichens form pale grey-brown to olive-green crusts on tree bark and are characterized by elongated, pencil-like slits containing spores, with distinctive tiny spines on internal filaments that help distinguish them from similar genera. The genus includes about 50 species found primarily in tropical and subtropical forests worldwide, where they grow on living tree bark and serve as indicators of relatively undisturbed woodland environments.
Calenia
Calenia is a genus of lichen-forming fungi within the family Gomphillaceae.
Hemithecium
Hemithecium is a genus of lichen-forming fungi in the family Graphidaceae. These lichens form pale grey-white to yellow-olive crusts on tree bark and are characterized by straight to weakly curved fruiting bodies with charcoal-black walls that create sharp contrasts against the surface, along with distinctive spine-tipped filaments lining their interior chambers. The genus has a pantropical distribution, growing on shaded tree trunks and large branches in primary or lightly disturbed evergreen forests, where their marked sensitivity to canopy opening makes them practical indicators of intact,
Platygramme
Platygramme is a genus of lichen-forming fungi in the family Graphidaceae consisting of about 27 species. The genus was circumscribed by Antoine Laurent Apollinaire Fée in 1874. The type species of the genus is P. caesiopruinosa. These bark-dwelling lichens are found in moist, shaded forests across tropical and warm temperate regions worldwide, where they form pale crusts that develop distinctive flattened, plate-like fruiting structures. A combination of several features characterizes Platygramme: the brown ascospores, a spore-producing layer (hymenium) speckled with granules, and elonga
Graphina
Graphina is a genus of script lichens in the family Graphidaceae. It has about 25 species. The genus was circumscribed in 1880 by Swiss lichenologist Johannes Müller Argoviensis. Müller Argoviensis did not indicate a type species for the genus in his original publication; David Hawksworth proposed to designate Graphina anguina as a lectotype in 1981.
Chroodiscus
Chroodiscus is a genus of leaf-dwelling lichens in the family Graphidaceae. These lichens form thin, smooth crusts directly on living leaves and are characterized by distinctive star-shaped fruiting bodies that split into triangular segments, with centres ranging from grey to bright scarlet-red depending on their chemical composition. The genus is found throughout tropical regions worldwide, growing in the shaded understory of rainforests from lowland areas to mountain cloud forests, where they serve as sensitive indicators of undisturbed forest conditions.
Aulaxina
Aulaxina is a genus of lichen-forming fungi in the family Gomphillaceae. Species of Aulaxina are found predominantly in humid tropical forests across Africa, Asia, and the Americas, where they grow on the surface of living leaves, a foliicolous lifestyle. The genus is recognised by its distinctive blackish fruiting bodies, which open in a star-like or slit-like pattern, and by the small dark hairs and club-shaped reproductive structures that dot the thallus surface. The exact number of species in the genus is uncertain, as different taxonomic databases currently disagree on which names are acc
Fissurina
Fissurina is a genus of lichenized fungi in the family Graphidaceae. It has about 160 species, most of which are found in tropical regions.
Caleniopsis
Caleniopsis is a genus of fungi in the family Gomphillaceae.
Stegobolus
Stegobolus is a genus of lichen-forming fungi in the family Graphidaceae. Established in 1845 by the French botanist Camille Montagne, the genus contains 16 species distinguished by their unique fruiting structures that initially appear drum-shaped and closed, later bursting open by shedding their roof-like covering. These bark-dwelling lichens are found in humid tropical and warm temperate forests worldwide, where they serve as indicators of undisturbed woodland habitats due to their sensitivity to canopy opening and drought.
Thalloloma
Thalloloma is a genus of lichen-forming fungi in the family Graphidaceae. Established in 1853 by the Italian botanist Vittore Trevisan de Saint-Léon, the genus comprises 20 species distinguished by their narrow script-like slits immersed in a pale crusty surface, often with dark brown or occasionally crimson-dusted fruiting structures. Found throughout tropical regions from sea level to mountain forests above 3,000 metres, these bark-dwelling lichens are more tolerant of light and moisture variation than many related species, allowing them to colonise forest edges and moderately sun-exposed lo
Diorygma
Diorygma is a genus of lichen-forming fungi in the family Graphidaceae. The genus was circumscribed by Franz Gerhard Eschweiler in 1824. Species of the genus are widely distributed in tropical and subtropical regions of the world. These lichens form paint-like crusts on bark and rock that range from chalky white to light green, with elongated, pencil-like slits containing their spores that may flex and branch across the surface. The genus was established in 1824 for tropical script lichens with large, many-celled spores, but molecular studies in the 2000s and 2010s expanded it significantly by
Diploschistella
Diploschistella is a genus of fungi in the family Gomphillaceae.
Pseudoramonia
Pseudoramonia is a genus of lichen-forming fungi in the family Graphidaceae. The genus was circumscribed by lichenologists Gintaras Kantvilas and Antonín Vězda in 2000, with the type species designated as Pseudoramonia stipitata.
Platythecium
Platythecium is a genus of lichen-forming fungi in the family Graphidaceae. It comprises 25 species. Described in 2002 by the German lichenologist Bettina Staiger, these bark-dwelling lichens are found in humid, shaded forests across tropical and warm temperate regions worldwide. They are characterized by their flattened, plate-like fruiting structures with completely blackened walls and are sensitive to forest disturbance, making their presence an indicator of long-established woodland conditions.
Actinoplaca
Actinoplaca is a small genus of lichen-forming fungi in the family Gomphillaceae. The genus is foliicolous, meaning its species grow on the surface of living leaves in humid tropical forests, and has been recorded from Central and South America and tropical Africa. The genus produces distinctive spherical structures on the thallus surface, now understood to be a type of asexual reproductive structure called '''', which were among the first such structures ever described in lichens.
Dyplolabia
Dyplolabia is a genus of lichen-forming fungi in the family Graphidaceae. These lichens form smooth, rather thick grey-yellow to olive-buff crusts on tree bark and are characterized by narrow, elongate fruiting bodies that are commonly hidden beneath a conspicuous white powdery coating. The genus has a pantropical distribution, growing on the smooth bark of trees and shrubs in both shaded rainforest understories and moderately exposed coastal woodlands, where they serve as indicators of long-established woodland habitat.
Fibrillithecis
Fibrillithecis is a genus of lichen-forming fungi in the family Graphidaceae. The genus was circumscribed in 2006 by the German lichenologist Andreas Frisch, with Fibrillithecis vernicosa assigned as the type species.
Anomomorpha
Anomomorpha is a genus of lichens in the family Graphidaceae. The genus, described in 1891, has a pantropical distribution. These lichens form pale grey to dull olive crusts on tree bark that often develop low wart-like bumps covered with powdery particles, and produce narrow, usually wavy slit-like fruiting bodies. They are found in tropical rainforests worldwide, growing on shaded to semi-exposed bark in primary or lightly disturbed evergreen forests, with some species being narrowly restricted to specific mountain cloud forests.
Aderkomyces
Aderkomyces is a genus of lichen-forming fungi in the family Gomphillaceae. Originally established in 1961 for a single species from Brazil, the genus was later expanded to include many species previously classified in the related genus Tricharia, all sharing distinctive white bristle-like structures and flattened fruiting bodies. Recent molecular studies suggest that Aderkomyces as currently defined may not represent a natural evolutionary group, leading to ongoing reclassification of many species into other genera.
Topeliopsis
Topeliopsis is a genus of lichen-forming fungi in the family Graphidaceae.
Leiorreuma
Leiorreuma is a genus of script lichens in the family Graphidaceae. It has 18 species. The genus was circumscribed by Franz Gerhard Eschweiler in 1824, with Leiorreuma hepaticum assigned as the type species.
Acanthotrema
Acanthotrema is a genus of lichens in the family Graphidaceae. The genus was circumscribed by German lichenologist Andreas Frisch in 2006, with Acanthotrema brasilianum assigned as the type species. Acanthotrema species are commonly found in rainforests ranging from lowland to montane environments.
Myriotrema
Myriotrema is a genus of lichen-forming fungi in the family Graphidaceae.
Tremotylium
Tremotylium is a genus of lichen-forming fungi in the family Graphidaceae. The genus was circumscribed by Finnish lichenologist William Nylander in 1865, but it was not published validly. He validated the genus and assigned T. angolense as the type species in 1868.
Q7076267
Ocellularia is a genus of lichens in the family Graphidaceae. The genus was circumscribed by Georg Friedrich Wilhelm Meyer in 1825.
Gyrotrema
Gyrotrema is a genus of lichen-forming fungi in the family Graphidaceae. There are six species assigned to the genus: Gyrotrema album Gyrotrema aurantiacum Gyrotrema flavum Gyrotrema papillatum Gyrotrema sinuosum Gyrotrema wirthii
Anomalographis
Anomalographis is a genus of two species of lichen-forming fungi in the family Graphidaceae. These lichens form pale grey to yellow-brown crusts and are characterized by elongated, slit-like fruiting bodies containing very small spores divided by a single internal wall. One species grows on shaded volcanic rock faces in Madeira, while the other was discovered growing on tree bases in northeastern Queensland rainforest, showing the genus can colonize both rock and bark surfaces in humid, shaded environments.
Ampliotrema
Ampliotrema is a genus of lichen-forming fungi in the family Graphidaceae. The genus was originally described invalidly in 2004, and validly two years later.
Pallidogramme
Pallidogramme is a genus of lichen-forming fungi in the family Graphidaceae. It has 8 species of corticolous (bark-dwelling), crustose lichens.
Glyphis
genus of fungi
Melanotrema
Melanotrema is a genus of lichen-forming fungi in the family Graphidaceae. The genus was circumscribed by German lichenologist Andreas Frisch in 2006, with M. platystomum assigned as the type species.
Gomphillus
Gomphillus is a genus of lichen-forming fungi in the family Gomphillaceae.
Corticifraga
Corticifraga is a genus of lichenicolous (lichen-dwelling) fungi in the family Gomphillaceae. The genus was established in 1990 and contains ten species that live as parasites on foliose lichens, mostly targeting species in the family Peltigeraceae. The produce no independent body (thallus) of their own, instead growing beneath the host's outer and eventually producing small, dark fruiting bodies that break through the surface.
Arthotheliopsis
Arthotheliopsis is a genus of fungi in the family Gomphillaceae. It comprises five species of crustose lichens. The genus was introduced by the Finnish lichenologist Edvard August Vainio in 1896, with A. hymenocarpoides assigned as the type species.
Redingeria
Redingeria is a genus of lichen-forming fungi in the family Graphidaceae. Established in 2006 and named after the Austrian botanist Karl Martin Redinger, the genus contains ten species that grow as bark-dwelling lichens in tropical forests worldwide. These lichens are distinguished by their script-like, elongated fruiting structures with thick black borders and are sensitive to forest disturbance, serving as indicators of undisturbed woodland habitats.
Thecaria
Thecaria is a genus of lichen-forming fungi in the family Graphidaceae.
Chapsa
Chapsa is a genus of lichens in the family Graphidaceae. These lichens form thin, grey-whitish to pale olive crusts on tree bark and are characterized by fruiting bodies that start as slits but expand into round to angular discs level with the surface, each bordered by a pale rim. The genus has a pantropical to warm-temperate distribution, growing on shaded bark in humid lowland or foothill rainforests, with over 60 species that often serve as indicators of undisturbed forest habitats.
Phaeotrema
Phaeotrema is a genus of lichen-forming fungi in the family Graphidaceae. The genus was circumscribed in 1887 by Johannes Müller Argoviensis, with P. subfarinosum assigned as the type species.
Leucodecton
Leucodecton is a genus of lichen-forming fungi in the family Graphidaceae. These lichens form thin, cream to pale brown crusts on bark or rock surfaces and reproduce through flask-shaped fruiting bodies that often appear in small, wart-like clusters. The genus currently includes more than 30 species found worldwide, with many recently described from tropical regions such as Sri Lanka and Costa Rica.
Melanotopelia
Melanotopelia is a genus of lichen-forming fungi in the family Graphidaceae. It has four species of corticolous (bark-dwelling), crustose lichens. This genus includes species characterised by dark pigmentation in their (a ring of tissue encircling their fruiting bodies), non-amyloid , and specific secondary metabolites.
Redonographa
Redonographa is a genus of lichen-forming fungi in the monogeneric family Redonographaceae. It has five species.
Cladosterigma
Cladosterigma is a fungal genus in the family Gomphillaceae. It is monospecific, comprising the single species Cladosterigma clavariella. The fungus is a hyperparasite: it grows on living leaves of myrtle relatives (genus Eugenia) where it parasitises a plant-pathogenic fungus in the genus Phyllachora. It is known only from an asexual state that forms conspicuous, yellow, branched synnemata (bundles of spore-bearing filaments) erupting through the leaf surface.
Wirthiotrema
Wirthiotrema is a genus of lichen-forming fungi in the family Graphidaceae. These tropical bark-dwelling lichens form yellow-green to light olive crusts with small, disc-like fruiting bodies that open through tiny circular pores. The genus contains six species found in lowland to mountain forests across tropical regions worldwide.