Pennsylvanian first appearances

Meganisoptera is an extinct order of large dragonfly-like insects, informally known as griffenflies or (incorrectly) as giant dragonflies. The order was formerly named Protodonata, the "proto-Odonata", for their similar appearance and supposed relation to modern Odonata (damselflies and dragonflies). They range in Palaeozoic (Late Carboniferous to Late Permian) times. Though most were only slightly larger than modern dragonflies, the order includes the largest known insect species, such as the late Carboniferous Meganeura monyi and the even larger early Permian Meganeuropsis permiana, with win

Sphenacodontidae (Greek: "wedge point tooth family") is an extinct family of sphenacodontoid synapsids. Small to large, advanced, carnivorous, Late Pennsylvanian to middle Permian "pelycosaurs". The most recent one, Dimetrodon angelensis, is from the latest Kungurian or, more likely, early Roadian San Angelo Formation. However, given the notorious incompleteness of the fossil record, a recent study concluded that the Sphenacodontidae may have become extinct as recently as the early Capitanian. Primitive forms were generally small (60 cm to 1 meter), but during the later part of the early

Parareptilia ("near-reptiles") is an extinct subclass of basal sauropsids ("reptiles"). Traditionally considered the sister taxon to Eureptilia (the group that likely contains all living reptiles and birds), its validity as a monophyletic clade has been disputed by modern cladistic analyses.

Caseasauria is one of the two main clades of early synapsids, the other being the Eupelycosauria. Caseasaurs are currently known only from the Late Carboniferous and the Permian, and include two superficially different families, the small insectivorous or carnivorous Eothyrididae, and the large, herbivorous Caseidae. These two groups share a number of specialised features associated with the morphology of the snout and external naris.

Eryops (; from Greek , , 'drawn-out' + , , 'face', because most of its skull was in front of its eyes) is a genus of extinct, amphibious temnospondyls. It contains the type species '''', the fossils of which are found mainly in early Permian deposits of the Texas Red Beds, and Eryops grandis from New Mexico. Fossils have also been found in late Carboniferous rocks from New Mexico and early Permian deposits of Oklahoma, Utah, the Pittsburgh tri-state region, and Prince Edward Island. Several complete skeletons of Eryops'' have been found in lower Permian rocks, but skull bones and teeth are its

Edaphosauridae is a family of mostly large (up to or more) Late Carboniferous to Early Permian synapsids. Edaphosaur fossils are so far known only from North America and Europe.

Varanopidae is an extinct family of amniotes known from the Late Carboniferous to Middle Permian that resembled monitor lizards (with the name of the group deriving from the monitor lizard genus Varanus) and may have filled a similar niche. Typically, they are considered to be relatively basal synapsids (and thus more closely related to mammals than to reptiles), although some studies from the late 2010s recovered them being taxonomically closer to diapsid reptiles; recent studies from the early 2020s support their traditional placement as synapsids on the basis of high degree of bone labyrint

The Palaeodictyoptera are an extinct order of medium-sized to very large, primitive Palaeozoic paleopterous insects. They are informative about the evolution of wings in insects.

Ophiacodontidae is an extinct family of early eupelycosaurs from the Carboniferous and Permian. Archaeothyris, and Clepsydrops were among the earliest ophiacodontids, appearing in the Late Carboniferous. Ophiacodontids are among the most basal synapsids, an offshoot of the lineage which includes therapsids and their descendants, the mammals. The group became extinct by the Kungurian or the Roadian, replaced by anomodonts, theriodonts, and the diapsid reptiles.

Araeoscelidia or Araeoscelida is a clade of extinct tetrapods (traditionally classified as diapsid reptiles) superficially resembling lizards, extending from the Late Carboniferous to the Early Permian. The group contains the genera Araeoscelis, Petrolacosaurus, the possibly aquatic Spinoaequalis, and less well-known genera such as Kadaliosaurus and Zarcasaurus. This clade has historically been considered to be the sister group to all (currently known) later diapsids, though studies from the early 2020s onwards have found it much closer to the base of Sauropsida unrelated to other diapsid rept

Seymouriamorpha were a small but widespread group of limbed vertebrates (tetrapods). They have long been considered stem-amniotes (reptiliomorphs), and most paleontologists still accept this point of view, but some analyses suggest that seymouriamorphs are stem-tetrapods (not more closely related to Amniota than to Lissamphibia).

Nectridea is an extinct order of lepospondyl tetrapods from the Carboniferous and Permian periods, including animals such as Diplocaulus. In appearance, they would have resembled modern newts or aquatic salamanders, although they are not close relatives of modern amphibians. They were characterized by long, flattened tails to aid in swimming, as well as numerous features of the vertebrae.

Microsauria is an extinct, possibly polyphyletic order of tetrapods from the late Carboniferous and early Permian periods. It is the most diverse and species-rich group of lepospondyls. Recently, Microsauria has been considered paraphyletic, as several other non-microsaur lepospondyl groups such as Lysorophia seem to be nested in it. Microsauria is now commonly used as a collective term for the grade of lepospondyls that were originally classified as members of Microsauria.

Captorhinidae is an extinct family of tetrapods, traditionally considered primitive reptiles, known from the late Carboniferous to the Late Permian. They had a cosmopolitan distribution across Pangea.

Diadectomorpha is a clade of large tetrapods that lived in Euramerica during the Carboniferous and Early Permian periods and in Asia during Late Permian (Wuchiapingian), They have typically been classified as advanced reptiliomorphs (transitional between "amphibians" sensu lato and amniotes) positioned close to, but outside of the clade Amniota, though some recent research has recovered them as the sister group to the traditional Synapsida within Amniota, based on inner ear anatomy and cladistic analyses. They include both large (up to 2 meters long) carnivorous and even larger (to 3 meters) h

Eryopidae are an extinct family of medium to large sized amphibious temnospondyls that lived from the latest Carboniferous (Pennsylvanian) to the early Permian period and inhabited North America and Europe. They were apex predators in the stream and lake habitats they inhabited. Their life cycle (exemplified by Onchiodon) consist of a small larvae that grows into a heavily ossified adult. This group are unusual in their variability in their morphology and development.

The Diaphanopterodea or Paramegasecoptera are an extinct order of moderate to large-sized Palaeozoic insects. They are first known from the Middle Carboniferous (late Serpukhovian or early Bashkirian in age), and include some of the earliest known flying insects.

Dissorophoidea is a clade of medium-sized, temnospondyl amphibians that appeared during the Moscovian in Euramerica, and continued through to the Late Permian and the Early Triassic of Gondwana. They are distinguished by various details of the skull, and many species seem to have been well adapted for life on land.

Dissorophidae is an extinct family of medium-sized temnospondyls that flourished during the late Carboniferous and early Permian periods. The clade is known almost exclusively from North America.

Diadectidae is an extinct family of early tetrapods that lived in what is now North America and Europe during the Late Carboniferous and Early Permian, and in Asia during the Late Permian. They were the first herbivorous tetrapods, and also the first fully terrestrial animals to attain large sizes. Footprints indicate that diadectids walked with an erect posture.

Megasecoptera is a Paleozoic insect order. There are 22 known families of megasecopterans, with about 35 known genera.

Eryopoidea is a clade of late Carboniferous and Permian temnospondyl amphibians, known from North America and Europe. Carroll (1998) includes no fewer than ten families, but Yates and Warren (2000) replaced this with a cladistic approach that includes three closely related families, the Eryopidae, Parioxyidae and Zatrachydidae. They define the Eryopoidea as all members of Euskelia in which the choana are relatively rounded and the iliac blade is vertical. A similar definition (without specifically naming Euskelia) is provided by Laurin and Steyer (2000).

Lysorophia is an order of fossorial Carboniferous and Permian tetrapods within the Recumbirostra. Lysorophians resembled small snakes, as their bodies are extremely elongate. There is a single family, the Molgophidae (previously known as Lysorophidae). Currently there are around five genera included within Lysorophia, although many may not be valid.

Zatracheidae (sometimes mistakenly spelled Zatrachydidae or Zatrachysdidae) is a family of Late Carboniferous and Early Permian temnospondyls known from North America and Europe. Zatracheidids are distinguished by lateral (sideways) bony protuberances of the quadratojugal bone of the skull, and a large opening in the snout called the internarial fontanelle (sometimes the internarial fenestra) that is bordered by enlarged premaxillae. The skull is flattened, with small orbits or eye sockets set far back. The opening in the snout may have housed a gland for producing a sticky substance so that p

Listracanthus is a genus of extinct chondrichthyan with uncertain affinities. Species of Listracanthus are known primarily from their tremendous, feather-like denticles, which range up to four inches in length. The denticles had a large main spine, from which secondary spines emanate from the sides, like the barbs of a feather or a comb. Listracanthus first appeared in late Carboniferous strata in North America, and eventually disappear from the fossil record some time during the Early Triassic.

The Paleodictyopterida or Palaeodictyopteroidea are an extinct superorder of Palaeozoic beaked insects, characterised by unique mouthparts consisting of 5 stylets. They represent the first important terrestrial herbivores, and the first major group of herbivorous insects. They appear during the Carboniferous (Serpukhovian stage) and continue through to the Late Permian. This large and diverse group includes 50% of all known Paleozoic insects. Palaeodictyopteroidea nymphs possessed movable wing pads and appear to have been able to perform simple flapping flight.

Euskelia is a clade of extinct temnospondyl amphibians. The naming derives from the ancient Greek eu, meaning "true", and skelos, meaning "limb", in reference to well-ossified limb bones with crests to which muscles were attached. Members of this group have the most ossified skeleton of all temnospondyls.

Archegosauroidea is an extinct superfamily of Permian temnospondyls. The superfamily is assigned to the clade Stereospondylomorpha and is the sister taxon to the suborder Stereospondyli. It includes the families Actinodontidae and Archegosauridae, and possibly the genus Intasuchus, which is placed within the monotypic family Intasuchidae. They were fully aquatic animals, and were metabolically and physiologically more similar to fish than modern amphibians.

Dvinosaurs are one of several new clades of temnospondyls named in the phylogenetic review of the group by Yates and Warren 2000. They represent a group of primitive semi-aquatic to completely aquatic temnospondyls, and are known from the Late Carboniferous to the Early Triassic, being most common in the Permian period. Their distinguishing characteristics are a reduction of the otic notch; the loss of a flange on the rear side of the pterygoid; and 28 or more presacral vertebrae.

Edopoidea is a clade of primitive temnospondyl amphibians including the genus Edops and the family Cochleosauridae. Edopoids are known from the Late Carboniferous and Early Permian of North America and Europe, and the Late Permian of Africa. They are among the most basal temnospondyls, and possess a number of primitive features that were lost in later members of the group.

Walchia is a primitive fossil conifer found in upper Pennsylvanian (Carboniferous) and lower Permian (about 310-290 Mya) rocks of Europe and North America. A forest of in-situ Walchia tree-stumps is located on the Northumberland Strait coast at Brule, Nova Scotia.

Aspidosaurus is an extinct genus of dissorophoid temnospondyl within the family Dissorophidae.

Limnoscelidae is a family of carnivorous diadectomorphs. They would have been the largest terrestrial carnivores of their day, the other large carnivores being aquatic or semi aquatic labyrinthodont amphibians. The Limnoscelidae themselves, being close to the ancestry of amniotes, would have been well adapted land animals, but still dependent on anamniote eggs, and possibly having a tadpole stage. Contrary to the more advanced herbivorous diadectids, the teeth retained labyrinthodont infolding of the enamel, and were pointed and slightly recurved at the tip.

Mazothairos (from Mazo, derived from its location of Mazon Creek and θαιρός, thairos, meaning 'hinge' in Greek) is an extinct genus of very large insect that lived during the Carboniferous period. It was a member of the order Palaeodictyoptera. Although it is only known from very fragmentary remains from a single fossil, it is estimated to have had a wingspan of around 56 centimeters, making it one of the largest-known insects, only being rivaled in size by the largest members of the order Meganisoptera, such as Meganeura and Meganeuropsis.

Gephyrostegidae is an extinct family of stegocephalian tetrapodomorphs from the Late Carboniferous including the genera Gephyrostegus, Bruktererpeton, and Eusauropleura. Gephyrostegus is from the Czech Republic, Brukterepeton is from Germany, and Eusauropleura is from the eastern United States.

Branchiosauridae is an extinct family of small amphibamiform temnospondyls with external gills and an overall juvenile appearance. The family has been characterized by hundreds of well-preserved specimens from the Permo-Carboniferous of Middle Europe. Specimens represent well defined ontogenetic stages and thus the taxon has been described to display paedomorphy (perennibranchiate). However, more recent work has revealed branchiosaurid taxa that display metamorphosing trajectories. The name Branchiosauridae (“Branchio” in Ancient Greek denoting gills and “saurus” meaning lizard) refers to the

Mycteroptidae or Mycteropidae are a family of eurypterids, a group of extinct chelicerate arthropods commonly known as "sea scorpions". The family is one of three families contained in the superfamily Mycteropoidea (along with Hibbertopteridae and Drepanopteridae), which in turn is one of four superfamilies classified as part of the suborder Stylonurina.

Limnarchia is a clade of temnospondyls. It includes the mostly Carboniferous-Permian age Dvinosauria and the mostly Permian-Triassic age Stereospondylomorpha. The clade was named in a 2000 phylogenetic analysis of stereospondyls and their relatives. Limnarchia means "lake rulers" in Greek, in reference to their aquatic lifestyles and long existence over a span of approximately 200 million years from the Late Carboniferous to the Early Cretaceous. In phylogenetic terms, Limnarchia is a stem-based taxon including all temnospondyls more closely related to Parotosuchus than to Eryops. It is the si

The Urocordylidae are an extinct family of nectridean lepospondyl amphibians. Urocordylids lived during the Late Carboniferous and Early Permian in what is now Europe and North America and are characterized by their very long, paddle-like tails. In life, they were probably newt-like and aquatic.

The Diplocaulidae ("double cauls") is an extinct family of "nectridean" tetrapodomorphs that arose during the Late Carboniferous and died out in the Late Permian. They are distinguished by the presence of strange, horn-like protrusions jutting out from the rear of their skulls; in some genera said protrusions gave their heads an almost boomerang-like outline. thumb|left|Diploceraspis burkei

Micromelerpetontidae (also spelled Micromelerpetidae) is an extinct family of dissorophoid temnospondyls that lived from the Late Carboniferous to the Early Permian in what is now Europe, with one Carboniferous species also known from North Africa. They were biologically similar to the related branchiosaurids, but proportionally akin to the unrelated microsaurs.

Arthrolycosa (meaning wolf [spider] with joints) is an extinct genus of arachnids, possibly spiders, that lived about 300-250 million years ago.

Oestocephalidae is an extinct family of Late Carboniferous aistopod tetrapodomorphs. Fossils have been found from Ohio, Illinois, and Colorado in the United States; England; and the Czech Republic. It includes the genera Coloraderpeton and Oestocephalus. Oestocephalids have robust skulls and narrow, rounded snouts. They possess heavily ossified gastralia and dorsal osteoderms. Like other aïstopods, oestocephalids were elongate, having approximately 110 vertebrae. Oestocephalidae was named in 2003, with the type species being Oestocephalus amphiuminus.

Gastrioceras is a goniatitid genus in the family Gastrioceratidae that lived during the late Carboniferous (Pennsylvanian and for most of the Permian.