microsporangium
thumb|Encephalartos villosus [[microsporophylls with microsporangia]] A microsporangium () is a sporangium that produces microspores that give rise to male gametophytes when they germinate. Microsporangia occur in all vascular plants that have heterosporic life cycles, such as seed plants, spike mosses and the aquatic fern genus Azolla. In gymnosperms and angiosperm anthers, the microsporangia produce microsporocytes, the microspore mother cells, which then produce four microspores through the process of meiosis. Microsporocytes are produced in the microsporangia of gymnosperm cones and the an
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thumb|Encephalartos villosus [[microsporophylls with microsporangia]] A microsporangium () is a sporangium that produces microspores that give rise to male gametophytes when they germinate. Microsporangia occur in all vascular plants that have heterosporic life cycles, such as seed plants, spike mosses and the aquatic fern genus Azolla. In gymnosperms and angiosperm anthers, the microsporangia produce microsporocytes, the microspore mother cells, which then produce four microspores through the process of meiosis. Microsporocytes are produced in the microsporangia of gymnosperm cones and the anthers of angiosperms. They are diploid microspore mother-cells, which then produce four haploid microspores by meiosis. These become pollen grains, within which the microspores divide twice by mitosis to produce a very simple gametophyte.
Heterosporous plants that produced microspores in microsporangia and megaspores in separate megasporangia evolved independently in several plant groups during the Devonian period. Fossils of these plants show that they produced endosporic gametophytes, meaning that their gametophytes were not free-living as in bryophytes but developed within the spores, as in modern heterosporic vascular plants.