Berothidae

The Berothidae are a family of winged insects of the order Neuroptera. They are known commonly as the beaded lacewings. The family was first named by Anton Handlirsch in 1906. The family consists of 24 genera and 110 living species distributed discontinuously worldwide, mostly in tropical and subtropical regions. Numerous extinct species have also been described. Their ecology is poorly known, but in the species where larval stages have been documented, the larvae are predators of termites.

==Systematics== thumb|Typical posture in life A considerable fossil diversity of beaded lacewings is known from the Late Jurassic onwards, containing numerous genera which are likewise basal or incertae sedis. Subfamily Berothimerobiinae Berothimerobius Subfamily Berothinae Asadeteva Barrowiella Berlekrumyia Berotha †Elektroberotha Eocene (Lutetian); Baltic amber, northern Europe Isoscelipteron Lekrugeria Lomamyia Nodalla Podallea Quasispermophorella Spermophorella Spiroberotha Stenobiella †Xenoberotha Green River Formation, United States, Eocene Subfamily Cyrenoberothinae †Aggregataberotha (Cenomanian, Burmese amber, Myanmar) †Araripeberotha Cretaceous (Aptian); Crato Formation, Brazil †Caririberotha Cretaceous (Aptian); Crato Formation, Brazil Cyrenoberotha Manselliberotha †Microberotha (Eocene (Ypresian), Hat Creek amber, Canada) Ormiscocerus †Protoberotha (Cenomanian, Burmese amber, Myanmar) †Sibelliberotha (Barremian, Lebanese amber) Speleoberotha Subfamily †Mesithoninae †Mesithone Callovian/Oxfordian; Karabastau Formation, Kazakhstan; Aptian; Zaza Formation, Turga Formation, Russia †Oloberotha Aptian; Yixian Formation, China Subfamily Nosybinae †Banoberotha Cretaceous (Barremian); Lebanese amber, Lebanon Naizema Nosybus Tanzanberotha Subfamily Nyrminae Nyrma Subfamily Protobiellinae Austroberothella Protobiella Subfamily Trichomatinae Trichoberotha Trichoma Subfamily Incertae sedis †Ansoberotha Cenomanian; Burmese amber, Myanmar †Berothone Callovian/Oxfordian; Karabastau Formation, Kazakhstan †Cantabroberotha Cretaceous (Albian); Spanish amber, Spain †Cornoberotha Cenomanian; Burmese amber, Myanmar †Dasyberotha Cenomanian; Burmese amber, Myanmar †Epimesoberotha Cretaceous (Berriasian); Lulworth Formation, United Kingdom †Ethiroberotha Cenomanian; Burmese amber, Myanmar †Haploberotha Cenomanian; Burmese amber, Myanmar †Iceloberotha Cenomanian; Burmese amber, Myanmar †Jersiberotha Cenomanian; Burmese amber, Myanmar; Cretaceous (Turonian); New Jersey amber, United States †Krokhathone Jurassic (Callovian/Oxfordian); Karabastau Formation, Kazakhstan †Maculaberotha Cenomanian; Burmese amber, Myanmar †Magniberotha Cenomanian; Burmese amber, Myanmar †Nascimberotha Turonian; New Jersey amber, United States †Plesiorobius Cretaceous (Campanian); Canadian amber, Canada; Campanian; Taymyr amber, Russia; Cenomanian, unknown formation, Russia †Proberotha Lutetian, Baltic Amber, northern Europe †Osmyloberotha Cenomanian; Burmese amber, Myanmar †Sinosmylites Daohugou, Haifanggou Formation, China, Callovian, Karabastau Formation, Kazakhstan, Callovian/Oxfordian, Ulaan-Ereg Formation, Mongolia, Late Jurassic (Tithonian) †Systenoberotha Cenomanian; Burmese amber, Myanmar †Telistoberotha Cenomanian; Burmese amber, Myanmar †Xiaoberotha Cenomanian; Burmese amber, Myanmar