Entelognathus primordialis

Entelognathus primordialis (“primordial complete jaw”) is an early placoderm from the late Silurian (Ludlow epoch) of Qujing, Yunnan, 419 million years ago. A team led by Min Zhu of the Academy of Sciences' Institute of Vertebrate Paleontology and Paleoanthropology in Beijing discovered the intact, articulated fossil in rock formations at Xiaoxiang reservoir.

==Specimen and taxonomy== left|thumb|Paratype specimen IVPP V32322 with scales and fin spines preserved The holotype of E. primordialis is the uncrushed and mostly intact anterior half of an individual with the articulating head and trunk armor preserved in three dimensions. The holotype is about long, and the live animal is estimated to have been over long. In overall form, the animal resembles primitive arthrodires, but the anatomy of the jaws strongly suggests the anatomies of bony fish and tetrapods. Specifically, this is the first stem gnathostome with dermal marginal jaw bones. These bones are the premaxilla, maxilla, and dentary. Most known placoderms had simple beak-like jaws made of bone plates. In 2023, newly described articulated specimen is shown, which have preserved large scales some of which are rhomboid, and fin spines on dorsal and anal fins (the latter of which was a trait that was previously only found in stem-group chondrichthyes).