Extant Cambrian first appearances

Vertebrates (), also called craniates, are animals with a vertebral column and a cranium. The vertebral column surrounds and protects the spinal cord, while the cranium protects the brain.

A chordate ( ) is a bilaterian animal belonging to the phylum Chordata ( ). All chordates possess, at some point during their larval or adult stages, five distinctive physical characteristics (synapomorphies) that distinguish them from other taxa: a notochord, a hollow dorsal nerve cord, an endostyle or thyroid, pharyngeal slits, and a post-anal tail.

thumb|Cornu aspersum (formerly Helix aspersa)—a common [[land snail]] thumb|Shell of Marisa cornuarietis, a [[freshwater snail]] Mollusca is a phylum of protostomic invertebrate animals, whose members are known as molluscs or mollusks (). 86,600 extant species of molluscs are recognized, making it the second-largest animal phylum after Arthropoda. The number of additional fossil species is estimated between 60,000 and 100,000, and the proportion of undescribed extant species is very high. Many taxa remain poorly studied.

Arthropods ( ) are invertebrates in the phylum Arthropoda. They possess an exoskeleton with a cuticle made of chitin, often mineralised with calcium carbonate, a body with differentiated (metameric) segments, and paired jointed appendages. In order to keep growing, they must go through stages of moulting, a process by which they shed their exoskeleton to reveal a new one. They form an extremely diverse group of up to ten million species.

thumb|Spotted jelly|Spotted jellies swimming in a Tokyo aquarium Jellyfish, also known as sea jellies or simply jellies, are the medusa-phase of certain gelatinous members of the subphylum Medusozoa, which is a major part of the phylum Cnidaria. Jellyfish are mainly free-swimming marine animals, although a few are anchored to the seabed by stalks rather than being motile. They are made of an umbrella-shaped main body made of mesoglea, known as the bell, and a collection of trailing tentacles on the underside.

Gastropods (; previously known as Univalves; class Gastropoda ) are a vast and diverse group of invertebrates within the phylum Mollusca, comprising the animals commonly known as snails and slugs. With an estimated 65,000 to 80,000 living species, they form the second-largest animal class after the insects. The fossil record of gastropods extends back to the Late Cambrian. , 721 families are recognized—476 extant (some with fossil representatives) and 245 extinct known only from fossils.

An echinoderm () is any animal of the phylum Echinodermata (), which includes starfish, brittle stars, sea urchins, sand dollars and sea cucumbers, as well as the sessile sea lilies or "stone lilies". While bilaterally symmetrical as larvae, as adults echinoderms are recognisable by their usually five-pointed radial symmetry (pentamerous symmetry), and are found on the sea bed at every ocean depth from the intertidal zone to the abyssal zone. The phylum contains about 7,600 living species, making it the second-largest group of deuterostomes after the chordates, as well as the largest marine-on

Crustaceans (from Latin word "crustacea" meaning: "those with shells" or "crusted ones") are mandibulate arthropods that are traditionally a part of the paraphyletic subphylum Crustacea (). Crustaceans are mainly aquatic, including decapods (shrimps, prawns, crabs, lobsters, and crayfish), seed shrimps, branchiopods, fish lice, krill, remipedes, isopods, barnacles, copepods, opossum shrimps, amphipods and mantis shrimp. It is now well accepted that hexapods (insects and entognathans) are cladistically crustaceans, with the two groups now combined in the monophyletic clade Pancrustacea. The thr

The annelids (), also known as the segmented worms, are animals that comprise the phylum Annelida (; ). The phylum contains over 22,000 extant species, including ragworms, earthworms, and leeches. The species exist in and have adapted to various ecologies – some in marine environments as distinct as tidal zones and hydrothermal vents, others in fresh water, and yet others in moist terrestrial environments.

Tardigrades (), also known as water bears or moss piglets, are a phylum of eight-legged segmented micro-animals. They were first described by the German zoologist Johann August Ephraim Goeze in 1773, who called them ' . In 1776, the Italian biologist Lazzaro Spallanzani named them Tardigrada', which means 'slow walkers'.

thumb|alt=Shell of the giant clam (Tridacna gigas)|Empty shell of the giant clam (Tridacna gigas) thumb|alt=Sword razor|Empty shells of the Ensis ensis|sword razor (Ensis ensis) Bivalvia () or bivalves, in previous centuries referred to as the Lamellibranchiata and Pelecypoda, is a class of aquatic molluscs (marine and freshwater) that have laterally compressed soft bodies enclosed by a calcified exoskeleton consisting of a hinged pair of half-shells known as valves. As a group, bivalves have no head and lack some typical molluscan organs such as the radula and the odontophore. Their gills hav

A cephalopod is any member of the molluscan class Cephalopoda (Greek plural , ; "head-feet") such as a squid, octopus, cuttlefish, or nautilus. These exclusively marine animals are characterized by bilateral body symmetry, a prominent head, and a set of arms or tentacles (muscular hydrostats) modified from the primitive molluscan foot. Fishers sometimes call cephalopods "inkfish", referring to their common ability to squirt ink. The study of cephalopods is a branch of malacology known as teuthology.

Ctenophora (; : ctenophore ) is a phylum of marine invertebrates, commonly known as comb jellies, that inhabit sea waters worldwide. They are notable for the groups of cilia they use for swimming (commonly referred to as "combs"), and they are the largest animals to swim with the help of cilia.

Tunicates are marine invertebrates belonging to the subphylum Tunicata ( ). This grouping is part of the Chordata, a phylum which includes all animals with dorsal nerve cords and notochords (including vertebrates). The subphylum was at one time called Urochordata, and the term urochordates is still sometimes used for these animals.

Hemichordata ( ) is a phylum which consists of triploblastic, eucoelomate, and bilaterally symmetrical marine deuterostome animals, generally considered the sister group of the echinoderms. They appear in the Lower or Middle Cambrian and include two main classes: Enteropneusta (acorn worms), and Pterobranchia. A third class, Planctosphaeroidea, is known only from the larva of a single species, Planctosphaera pelagica. The class Graptolithina, formerly considered extinct, is now placed within the pterobranchs, represented by a single living genus Rhabdopleura.

Agnatha (; ) or jawless fish is a paraphyletic infraphylum of animals in the subphylum Vertebrata of the phylum Chordata, characterized by the lack of jaws. The group consists of both living (cyclostomes such as hagfishes and lampreys) and extinct clades (e.g. conodonts and cephalaspidomorphs, among others). They are sister to vertebrates with jaws known as gnathostomes, who evolved from jawless ancestors during the early Silurian by developing folding articulations in the first pairs of gill arches.

Brachiopods (), phylum Brachiopoda, are a phylum of animals that have hard "valves" (shells) on the upper and lower surfaces, unlike the left and right arrangement in bivalve molluscs. Brachiopod valves are hinged at the rear end, while the front can be opened for feeding or closed for protection.

Malacostraca is the second largest of the six classes of pancrustaceans behind insects, containing about 40,000 living species, divided among 16 orders. Its members, the malacostracans, display a great diversity of body forms and include crabs, lobsters, spiny lobsters, crayfish, shrimp, krill, prawns, isopods, amphipods, mantis shrimp, and many other less familiar animals. They are abundant in all marine environments and have colonised freshwater and terrestrial habitats. They are segmented animals, united by a common body plan comprising 20 body segments (rarely 21), and divided into a head,

Branchiopoda (from Ancient Greek βράγχια (bránkhia), meaning "gill", and πούς (poús), meaning "foot") is a class of crustaceans. It comprises fairy shrimp, clam shrimp, Diplostraca (or Cladocera), Notostraca, the Devonian Lepidocaris and possibly the Cambrian Rehbachiella. They are mostly small, freshwater animals that feed on plankton and detritus.

class of marine arthropods

Polychaeta () is a paraphyletic class of generally marine annelid worms, commonly called bristle worms or polychaetes (). Each body segment has a pair of fleshy protrusions called parapodia which bear many chitinous bristles called chaetae, hence their name.

Foraminifera ( ; Latin for "hole bearers"; informally called "forams") are single-celled organisms, members of a phylum or class of Rhizarian protists characterized by streaming granular ectoplasm for catching food and other uses; and commonly an external shell called a test of diverse forms and materials. Tests of chitin (found in some simple genera, and Textularia in particular) are believed to be the most primitive type. Most foraminifera are marine, the majority of which live on or within the seafloor sediment (i.e., are benthic, with different sized species playing a role within the macro

The Chaetognatha or chaetognaths (meaning bristle-jaws) are a phylum of predatory marine worms that are a major component of plankton worldwide. Commonly known as arrow worms, they are mostly pelagic; however about 20% of the known species are benthic, and can attach to algae and rocks. They are found in all marine waters, from surface tropical waters and shallow tide pools to the deep sea and polar regions. Most chaetognaths are transparent and are torpedo shaped, but some deep-sea species are orange. They range in size from .

Entoprocta (), or Kamptozoa , commonly called goblet worms, is a phylum of mostly-sessile aquatic animals, ranging from long. Mature individuals are goblet-shaped, on relatively long stalks. They have a "crown" of solid tentacles whose cilia generate water currents that draw food particles towards the mouth, and both the mouth and anus lie inside the "crown". The superficially similar Bryozoa (Ectoprocta) have the anus outside a "crown" of hollow tentacles. Most families of entoprocts are colonial, and all but 2 of the 150 species are marine. A few solitary species can move slowly.

Ecdysozoa () is a group of protostome animals, including Arthropoda (insects, chelicerates (including arachnids), crustaceans, and myriapods), Nematoda, and several smaller phyla. The grouping of these animal phyla into a single clade was first proposed by Eernisse et al. (1992) based on a phylogenetic analysis of 141 morphological characters of ultrastructural and embryological phenotypes. This clade, that is, a group consisting of a common ancestor and all its descendants, was formally named by Aguinaldo et al. in 1997, based mainly on phylogenetic trees constructed using 18S ribosomal RNA g

Ascidiacea, commonly known as the ascidians or sea squirts, is a paraphyletic class in the subphylum Tunicata of sac-like marine invertebrate filter feeders. Ascidians are characterized by a tough outer test or "tunic" made of the polysaccharide cellulose.

The Sipuncula or Sipunculida (common names sipunculid worms or peanut worms) is a class containing about 162 species of marine annelid worms, that have secondarily lost their segmentation. Sipuncula was once considered a phylum of unsegmented worms, but was demoted to a class of Annelida, based on recent molecular work.

The Radiolaria, also called Radiozoa and informally called radiolarians, are unicellular eukaryotes of diameter 0.1–0.2 mm that produce intricate mineral skeletons, typically with a central capsule dividing the cell into the inner and outer portions of endoplasm and ectoplasm. The elaborate mineral skeleton is usually made of silica. They are found as zooplankton throughout the global ocean. As zooplankton, radiolarians are primarily heterotrophic, but many have photosynthetic endosymbionts and are, therefore, considered mixotrophs. The skeletal remains of some types of radiolarians make

thumb|Shell of Monoplacophora Monoplacophora , meaning "bearing one plate", is a polyphyletic class of molluscs with a cap-like shell, inhabiting deep sea environments. Extant representatives were not recognized as such until 1952; previously they were known only from the fossil record, and were thought to have become extinct 375 million years ago.

Loricifera (from Latin, lorica, corselet (armour) + ferre, to bear), also known as corset animals, is a phylum of very small to microscopic marine scalidophoran sediment-dwelling animals with 43 described species and approximately 100 more that have been collected and not yet described. Their sizes range from 100 μm to .

Lophotrochozoa (, "crest/wheel animals") is a clade of protostome animals within the Spiralia. The taxon was established as a monophyletic group based on molecular evidence. The clade includes animals like annelids, molluscs, bryozoans, and brachiopods.

thumb|Bolosoma stalked glass sponge Hexactinellid sponges are sponges with a skeleton made of four- and/or six-pointed siliceous spicules, often referred to as glass sponges. They are usually classified along with other sponges in the phylum Porifera, but some researchers consider them sufficiently distinct to deserve their own phylum, Symplasma. Some experts believe that glass sponges are the longest-lived animals on earth; these scientists tentatively estimate a maximum age of up to 15,000 years.

Nautiloids are a group of cephalopods (Mollusca) which originated in the Late Cambrian and are represented today by the living Nautilus and Allonautilus. Fossil nautiloids are diverse and species rich, with over 2,500 recorded species. They flourished during the early Paleozoic era, when they constituted the main predatory animals. Early in their evolution, nautiloids developed an extraordinary diversity of shell shapes, including coiled morphologies and giant straight-shelled forms (orthocones). No orthoconic and only a handful of coiled species, the nautiluses, survive to the present day.

The Pentastomida are an enigmatic group of parasitic arthropods commonly known as tongue worms due to the resemblance of the species of the genus Linguatula to a vertebrate tongue; molecular studies point to them being highly derived crustaceans.

Panarthropoda is a clade comprising the greatest diversity of animal groups. It contains the extant phyla Arthropoda (Euarthropoda), Tardigrada (water bears) and Onychophora (velvet worms), although the relationships among these remained uncertain according to studies published in 2023 and 2024. Panarthropods also include extinct marine legged worms known as lobopodians ("Lobopodia"), a paraphyletic group where the last common ancestor and basal members (stem-group) of each extant panarthropod phylum are thought to have risen. However the term "Lobopodia" is sometimes expanded to include tardi

order of cnidarians

Medusozoa is a clade in the phylum Cnidaria, and is often considered a subphylum. It includes the classes Hydrozoa, Scyphozoa, Staurozoa and Cubozoa. Medusozoans are distinguished by having a medusa stage in their often complex life cycle, a medusa typically being an umbrella-shaped body with stinging tentacles around the edge. With the exception of some Hydrozoa (and Polypodiozoa), all are called jellyfish in their free-swimming medusa phase. ==Naming== The Medusozoa are named after Medusa from ancient Greek mythology.

Phyllocarida is a subclass of crustaceans, comprising the extant order Leptostraca and the extinct orders Hymenostraca and Archaeostraca. This clade of marine crustaceans diversified extensively during the Ordovician. left|thumb|Undescribed fossil of a very large phyllocarid from the Yukon ([[Devonian)]]

Crinozoa is a subphylum of mostly sessile echinoderms, of which the crinoids, or sea lilies and feather stars, are the only extant members. Crinozoans have an extremely extensive fossil history.

Eleutherozoa is a subphylum of echinoderms. They are mobile animals with the mouth directed towards the substrate. They usually have a madreporite, tube feet, and moveable spines of some sort. It includes all living echinoderms except for crinoids. The monophyly of Eleutherozoa has been proven sufficiently well to be considered "uncontroversial."

Lingulata is a class of brachiopods, among the oldest of all brachiopods having existed since the Cambrian period (). They are also among the most morphologically conservative of the brachiopods, having lasted from their earliest appearance to the present with very little change in shape. Shells of living specimens found today in the waters around Japan are almost identical to ancient Cambrian fossils.

Archaeogastropoda (also known as Aspidobranchia) was a taxonomic order of snails used in older classifications of gastropods, i.e. snails and slugs. Archeogastropoda are primarily marine prosobranch gastropod mollusks, mainly herbivores, typically having two gills and a double-chambered heart, with the eggs and sperm discharged directly into the water, excepting terrestrial species. They were traditionally regarded as a relatively primitive group.

Lingulida is an order of brachiopods.

Rhabdopleura is a genus of colonial sessile hemichordates belonging to the Pterobranchia class. They are exclusively marine, benthic organisms whose species occur within all major oceans and range in habitat from intertidal to . As one of the oldest living genera with a fossil record dating back to the Middle Cambrian, it is also considered to be the only living genus of graptolites.

Oligostraca is a superclass of crustaceans. It consists of the following classes:

Allogromiida is an order of single-chambered, mostly organic-walled foraminiferans, including some that produce agglutinated tests (Lagynacea). Genetic studies indicate that some foraminiferans with agglutinated tests, previously included in the Textulariida or as their own order Astrorhizida, may also belong here. Allogromiids produce relatively simple tests, usually with a single chamber, similar to those of other protists such as Gromia. They are found as both marine and freshwater forms, and are the oldest forms known from the fossil record.

The Textulariida are an order of foraminifera that produce agglutinated shells or tests. An agglutinated test is one made of foreign particles glued together with an organic or calcareous cement to form an external shell on the outside of the organism. Commonly, the order had been made up of all species of Foraminifera with these types of shells, but genetic studies indicate these organisms do not form an evolutionary group, and several superfamilies in the order have been moved to the order Allogromiida. The remaining forms are sometimes divided into three orders: the Trochamminida and Lituol